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BioLiP
Download all results in tab-seperated text for 832530 receptor-ligand interactions, whose format is explained at readme.txt.
  • Hover over PDB to view the title of the structure. Click PDB to view the structure at the RCSB PDB database. Resolution -1.00 means the resolution is unavailable, e.g., for NMR structures.
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  • Hover over Ligand to view the full ligand name. Click Ligand to view the 2D diagram and other detail information of the ligand.
  • Hover over EC number to view the full name of enzymatic activity.
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    # PDB
    (Resolution Å)
    Site # Ligand EC number GO terms UniProt PubMed Binding
    affinity
    15401 1go7:P (2.1) BS02 CA 3.4.24.40 GO:0004222 ... P16317 11445573
    15402 1go7:P (2.1) BS03 CA 3.4.24.40 GO:0004222 ... P16317 11445573
    15403 1go7:P (2.1) BS04 CA 3.4.24.40 GO:0004222 ... P16317 11445573
    15404 1go7:P (2.1) BS05 CA 3.4.24.40 GO:0004222 ... P16317 11445573
    15405 1go7:P (2.1) BS06 ZN 3.4.24.40 GO:0004222 ... P16317 11445573
    15406 1go7:P (2.1) BS07 CA 3.4.24.40 GO:0004222 ... P16317 11445573
    15407 1go7:P (2.1) BS08 CA 3.4.24.40 GO:0004222 ... P16317 11445573
    15408 1go8:P (2.0) BS01 CA 3.4.24.40 GO:0004222 ... P16317 11445573
    15409 1go8:P (2.0) BS02 CA 3.4.24.40 GO:0004222 ... P16317 11445573
    15410 1go8:P (2.0) BS03 CA 3.4.24.40 GO:0004222 ... P16317 11445573
    15411 1go8:P (2.0) BS04 CA 3.4.24.40 GO:0004222 ... P16317 11445573
    15412 1go8:P (2.0) BS05 CA 3.4.24.40 GO:0004222 ... P16317 11445573
    15413 1go8:P (2.0) BS06 CA 3.4.24.40 GO:0004222 ... P16317 11445573
    15414 1go8:P (2.0) BS07 ZN 3.4.24.40 GO:0004222 ... P16317 11445573
    15415 1go8:P (2.0) BS08 CA 3.4.24.40 GO:0004222 ... P16317 11445573
    15416 1gof:A (1.7) BS01 CU 1.1.3.9 GO:0005576 ... P0CS93 2002850
    15417 1gog:A (1.9) BS01 CU 1.1.3.9 GO:0005576 ... P0CS93 2002850
    15418 1goj:A (2.3) BS01 ADP 3.6.4.4 GO:0003777 ... P48467 11707393
    15419 1gol:A (2.8) BS01 MG 2.7.11.24 GO:0000165 ... P63086 8639522
    15420 1gol:A (2.8) BS02 ATP 2.7.11.24 GO:0000165 ... P63086 8639522
    15421 1gon:A (2.2) BS01 SO4 3.2.1.21 GO:0004553 ... Q59976 N/A
    15422 1goq:A (1.8) BS01 XYP 3.2.1.8 GO:0004553 ... P23360 11741607
    15423 1goq:A (1.8) BS02 XYP 3.2.1.8 GO:0004553 ... P23360 11741607
    15424 1goq:A (1.8) BS03 XYP 3.2.1.8 GO:0004553 ... P23360 11741607
    15425 1goq:A (1.8) BS04 XYP 3.2.1.8 GO:0004553 ... P23360 11741607
    15426 1goq:A (1.8) BS05 XYP 3.2.1.8 GO:0004553 ... P23360 11741607
    15427 1gor:A (1.7) BS01 XYS 3.2.1.8 GO:0004553 ... P23360 11741607
    15428 1gor:A (1.7) BS02 XYP 3.2.1.8 GO:0004553 ... P23360 11741607
    15429 1gos:A (3.0) BS01 FAD 1.4.3.21
    1.4.3.4
    GO:0005515 ... P27338 11753429
    15430 1gos:A (3.0) BS02 NYP 1.4.3.21
    1.4.3.4
    GO:0005515 ... P27338 11753429
    15431 1gos:B (3.0) BS01 FAD 1.4.3.21
    1.4.3.4
    GO:0005515 ... P27338 11753429
    15432 1gos:B (3.0) BS02 NYP 1.4.3.21
    1.4.3.4
    GO:0005515 ... P27338 11753429
    15433 1got:A (2.0) BS01 GDP ? GO:0000035 ... P04695 8552184
    15434 1gov:A (2.0) BS01 SO4 3.1.27.- GO:0003723 ... P00649 11976484
    15435 1gov:A (2.0) BS02 SO4 3.1.27.- GO:0003723 ... P00649 11976484
    15436 1gov:B (2.0) BS01 SO4 3.1.27.- GO:0003723 ... P00649 11976484
    15437 1gov:B (2.0) BS02 SO4 3.1.27.- GO:0003723 ... P00649 11976484
    15438 1gox:A (2.0) BS01 FMN 1.1.3.15 GO:0003973 ... P05414 2681790
    15439 1goy:A (2.0) BS01 3GP 3.1.27.- GO:0003723 ... P00649 11976484
    15440 1goy:A (2.0) BS02 SO4 3.1.27.- GO:0003723 ... P00649 11976484
    15441 1goy:B (2.0) BS01 SO4 3.1.27.- GO:0003723 ... P00649 11976484
    15442 1goy:B (2.0) BS02 SO4 3.1.27.- GO:0003723 ... P00649 11976484
    15443 1gp2:A (2.3) BS01 GDP ? GO:0000287 ... P10824 8521505
    15444 1gp4:A (2.1) BS01 AKG 1.14.20.4 GO:0007033 ... Q96323 11796114
    15445 1gp5:A (2.2) BS01 AKG 1.14.20.4 GO:0007033 ... Q96323 11796114
    15446 1gp5:A (2.2) BS02 DQH 1.14.20.4 GO:0007033 ... Q96323 11796114
    15447 1gp5:A (2.2) BS03 DH2 1.14.20.4 GO:0007033 ... Q96323 11796114
    15448 1gp5:A (2.2) BS04 FE 1.14.20.4 GO:0007033 ... Q96323 11796114
    15449 1gp6:A (1.75) BS01 QUE 1.14.20.4 GO:0007033 ... Q96323 11796114
    15450 1gp6:A (1.75) BS02 DH2 1.14.20.4 GO:0007033 ... Q96323 11796114
    15451 1gp6:A (1.75) BS03 FE2 1.14.20.4 GO:0007033 ... Q96323 11796114
    15452 1gp7:A (2.6) BS01 CA 3.1.1.4 GO:0004623 ... P80966 12217659
    15453 1gp7:B (2.6) BS01 CA 3.1.1.4 GO:0004623 ... P80966 12217659
    15454 1gp7:C (2.6) BS01 CA 3.1.1.4 GO:0004623 ... P80966 12217659
    15455 1gpa:A (2.9) BS01 PLP 2.4.1.1 GO:0000166 ... P00489 1900534
    15456 1gpa:B (2.9) BS01 PLP 2.4.1.1 GO:0000166 ... P00489 1900534
    15457 1gpa:C (2.9) BS01 PLP 2.4.1.1 GO:0000166 ... P00489 1900534
    15458 1gpa:D (2.9) BS01 PLP 2.4.1.1 GO:0000166 ... P00489 1900534
    15459 1gpb:A (1.9) BS01 PLP 2.4.1.1 GO:0000166 ... P00489 N/A
    15460 1gpc:A (2.2) BS01 ZN ? GO:0003697 ... P03695 7630406
    15461 1gpd:G (2.9) BS01 NAD 1.2.1.12 GO:0004365 ... P00357 127793
    15462 1gpd:R (2.9) BS01 PO4 1.2.1.12 GO:0004365 ... P00357 127793
    15463 1gpd:R (2.9) BS02 NAD 1.2.1.12 GO:0004365 ... P00357 127793
    15464 1gpe:A (1.8) BS01 BMA 1.1.3.4 GO:0005576 ... P81156 10216293
    15465 1gpe:A (1.8) BS02 MAN 1.1.3.4 GO:0005576 ... P81156 10216293
    15466 1gpe:A (1.8) BS03 FAD 1.1.3.4 GO:0005576 ... P81156 10216293
    15467 1gpe:B (1.8) BS01 BMA 1.1.3.4 GO:0005576 ... P81156 10216293
    15468 1gpe:B (1.8) BS02 MAN 1.1.3.4 GO:0005576 ... P81156 10216293
    15469 1gpe:B (1.8) BS03 FAD 1.1.3.4 GO:0005576 ... P81156 10216293
    15470 1gph:1 (3.0) BS01 SF4 2.4.2.14 GO:0000287 ... P00497 8197456
    15471 1gph:1 (3.0) BS02 AMP 2.4.2.14 GO:0000287 ... P00497 8197456
    15472 1gph:1 (3.0) BS03 AMP 2.4.2.14 GO:0000287 ... P00497 8197456
    15473 1gph:1 (3.0) BS04 AMP 2.4.2.14 GO:0000287 ... P00497 8197456
    15474 1gph:2 (3.0) BS01 AMP 2.4.2.14 GO:0000287 ... P00497 8197456
    15475 1gph:2 (3.0) BS02 SF4 2.4.2.14 GO:0000287 ... P00497 8197456
    15476 1gph:2 (3.0) BS03 AMP 2.4.2.14 GO:0000287 ... P00497 8197456
    15477 1gph:2 (3.0) BS04 AMP 2.4.2.14 GO:0000287 ... P00497 8197456
    15478 1gph:3 (3.0) BS01 SF4 2.4.2.14 GO:0000287 ... P00497 8197456
    15479 1gph:3 (3.0) BS02 AMP 2.4.2.14 GO:0000287 ... P00497 8197456
    15480 1gph:3 (3.0) BS03 AMP 2.4.2.14 GO:0000287 ... P00497 8197456
    15481 1gph:3 (3.0) BS04 AMP 2.4.2.14 GO:0000287 ... P00497 8197456
    15482 1gph:4 (3.0) BS01 SF4 2.4.2.14 GO:0000287 ... P00497 8197456
    15483 1gph:4 (3.0) BS02 AMP 2.4.2.14 GO:0000287 ... P00497 8197456
    15484 1gph:4 (3.0) BS03 AMP 2.4.2.14 GO:0000287 ... P00497 8197456
    15485 1gpj:A (1.95) BS01 GLU 1.2.1.70 GO:0006779 ... Q9UXR8 11726494
    15486 1gpj:A (1.95) BS02 GMC 1.2.1.70 GO:0006779 ... Q9UXR8 11726494
    15487 1gpk:A (2.1) BS01 HUP 3.1.1.7 GO:0001507 ... P04058 12196020 MOAD: Ki=4.3uM
    PDBbind: -logKd/Ki=5.37, Ki=4.3uM
    BindingDB: IC50=11.4nM
    15488 1gpl:A (2.01) BS01 CA 3.1.1.26
    3.1.1.3
    GO:0004806 ... P16233
    P81139
    8939760
    15489 1gpm:A (2.2) BS01 POP 6.3.5.2 GO:0003921 ... P04079 8548458
    15490 1gpm:A (2.2) BS02 AMP 6.3.5.2 GO:0003921 ... P04079 8548458
    15491 1gpm:B (2.2) BS01 POP 6.3.5.2 GO:0003921 ... P04079 8548458
    15492 1gpm:B (2.2) BS02 AMP 6.3.5.2 GO:0003921 ... P04079 8548458
    15493 1gpm:C (2.2) BS01 POP 6.3.5.2 GO:0003921 ... P04079 8548458
    15494 1gpm:C (2.2) BS02 AMP 6.3.5.2 GO:0003921 ... P04079 8548458
    15495 1gpm:D (2.2) BS01 POP 6.3.5.2 GO:0003921 ... P04079 8548458
    15496 1gpm:D (2.2) BS02 AMP 6.3.5.2 GO:0003921 ... P04079 8548458
    15497 1gpn:A (2.35) BS01 HUB 3.1.1.7 GO:0001507 ... P04058 12196020 MOAD: Ki=0.334uM
    PDBbind: -logKd/Ki=6.48, Ki=0.334uM
    BindingDB: Ki=334nM
    15498 1gpu:A (1.86) BS01 CA 2.2.1.1 GO:0003824 ... P23254 11773632
    15499 1gpu:A (1.86) BS02 THD 2.2.1.1 GO:0003824 ... P23254 11773632
    15500 1gpu:A (1.86) BS03 THD 2.2.1.1 GO:0003824 ... P23254 11773632
    15501 1gpu:B (1.86) BS01 THD 2.2.1.1 GO:0003824 ... P23254 11773632
    15502 1gpu:B (1.86) BS02 CA 2.2.1.1 GO:0003824 ... P23254 11773632
    15503 1gpu:B (1.86) BS03 THD 2.2.1.1 GO:0003824 ... P23254 11773632
    15504 1gpw:C (2.4) BS01 PO4 4.3.2.10 GO:0000105 ... Q9X0C6 11839304
    15505 1gpw:C (2.4) BS02 PO4 4.3.2.10 GO:0000105 ... Q9X0C6 11839304
    15506 1gpw:E (2.4) BS01 PO4 4.3.2.10 GO:0000105 ... Q9X0C6 11839304
    15507 1gpx:A (-1.00) BS01 GA ? GO:0005515 ... P00259 9835048
    15508 1gpy:A (2.4) BS01 G6P 2.4.1.1 GO:0000166 ... P00489 8331662 MOAD: Kd~25uM
    PDBbind: -logKd/Ki=4.70, Kd=20uM
    15509 1gpy:A (2.4) BS02 PLP 2.4.1.1 GO:0000166 ... P00489 8331662
    15510 1gpz:A (2.9) BS01 FUC 3.4.21.41 GO:0004252 ... P00736 11823416
    15511 1gpz:A (2.9) BS02 FUC 3.4.21.41 GO:0004252 ... P00736 11823416
    15512 1gpz:B (2.9) BS01 FUC 3.4.21.41 GO:0004252 ... P00736 11823416
    15513 1gq1:A (1.4) BS01 HEC 1.7.2.1
    1.7.99.1
    GO:0009055 ... P72181 12556530
    15514 1gq1:A (1.4) BS02 DHE 1.7.2.1
    1.7.99.1
    GO:0009055 ... P72181 12556530
    15515 1gq1:A (1.4) BS03 SO4 1.7.2.1
    1.7.99.1
    GO:0009055 ... P72181 12556530
    15516 1gq1:A (1.4) BS04 SO4 1.7.2.1
    1.7.99.1
    GO:0009055 ... P72181 12556530
    15517 1gq1:A (1.4) BS05 SO4 1.7.2.1
    1.7.99.1
    GO:0009055 ... P72181 12556530
    15518 1gq1:B (1.4) BS01 HEC 1.7.2.1
    1.7.99.1
    GO:0009055 ... P72181 12556530
    15519 1gq1:B (1.4) BS02 DHE 1.7.2.1
    1.7.99.1
    GO:0009055 ... P72181 12556530
    15520 1gq1:B (1.4) BS03 SO4 1.7.2.1
    1.7.99.1
    GO:0009055 ... P72181 12556530
    15521 1gq1:B (1.4) BS04 SO4 1.7.2.1
    1.7.99.1
    GO:0009055 ... P72181 12556530
    15522 1gq1:B (1.4) BS05 SO4 1.7.2.1
    1.7.99.1
    GO:0009055 ... P72181 12556530
    15523 1gq1:B (1.4) BS06 SO4 1.7.2.1
    1.7.99.1
    GO:0009055 ... P72181 12556530
    15524 1gq1:B (1.4) BS07 SO4 1.7.2.1
    1.7.99.1
    GO:0009055 ... P72181 12556530
    15525 1gq2:A (2.5) BS01 NAP 1.1.1.40 GO:0004470 ... P40927 11790843
    15526 1gq2:A (2.5) BS02 OXL 1.1.1.40 GO:0004470 ... P40927 11790843
    15527 1gq2:A (2.5) BS03 MN 1.1.1.40 GO:0004470 ... P40927 11790843
    15528 1gq2:B (2.5) BS01 NAP 1.1.1.40 GO:0004470 ... P40927 11790843
    15529 1gq2:B (2.5) BS02 OXL 1.1.1.40 GO:0004470 ... P40927 11790843
    15530 1gq2:B (2.5) BS03 MN 1.1.1.40 GO:0004470 ... P40927 11790843
    15531 1gq2:C (2.5) BS01 NAP 1.1.1.40 GO:0004470 ... P40927 11790843
    15532 1gq2:C (2.5) BS02 OXL 1.1.1.40 GO:0004470 ... P40927 11790843
    15533 1gq2:C (2.5) BS03 MN 1.1.1.40 GO:0004470 ... P40927 11790843
    15534 1gq2:D (2.5) BS01 NAP 1.1.1.40 GO:0004470 ... P40927 11790843
    15535 1gq2:D (2.5) BS02 OXL 1.1.1.40 GO:0004470 ... P40927 11790843
    15536 1gq2:D (2.5) BS03 MN 1.1.1.40 GO:0004470 ... P40927 11790843
    15537 1gq2:E (2.5) BS01 NAP 1.1.1.40 GO:0004470 ... P40927 11790843
    15538 1gq2:E (2.5) BS02 OXL 1.1.1.40 GO:0004470 ... P40927 11790843
    15539 1gq2:E (2.5) BS03 MN 1.1.1.40 GO:0004470 ... P40927 11790843
    15540 1gq2:F (2.5) BS01 NAP 1.1.1.40 GO:0004470 ... P40927 11790843
    15541 1gq2:F (2.5) BS02 OXL 1.1.1.40 GO:0004470 ... P40927 11790843
    15542 1gq2:F (2.5) BS03 MN 1.1.1.40 GO:0004470 ... P40927 11790843
    15543 1gq2:G (2.5) BS01 NAP 1.1.1.40 GO:0004470 ... P40927 11790843
    15544 1gq2:G (2.5) BS02 OXL 1.1.1.40 GO:0004470 ... P40927 11790843
    15545 1gq2:G (2.5) BS03 MN 1.1.1.40 GO:0004470 ... P40927 11790843
    15546 1gq2:H (2.5) BS01 NAP 1.1.1.40 GO:0004470 ... P40927 11790843
    15547 1gq2:H (2.5) BS02 OXL 1.1.1.40 GO:0004470 ... P40927 11790843
    15548 1gq2:H (2.5) BS03 MN 1.1.1.40 GO:0004470 ... P40927 11790843
    15549 1gq2:I (2.5) BS01 NAP 1.1.1.40 GO:0004470 ... P40927 11790843
    15550 1gq2:I (2.5) BS02 OXL 1.1.1.40 GO:0004470 ... P40927 11790843
    15551 1gq2:I (2.5) BS03 MN 1.1.1.40 GO:0004470 ... P40927 11790843
    15552 1gq2:J (2.5) BS01 NAP 1.1.1.40 GO:0004470 ... P40927 11790843
    15553 1gq2:J (2.5) BS02 OXL 1.1.1.40 GO:0004470 ... P40927 11790843
    15554 1gq2:J (2.5) BS03 MN 1.1.1.40 GO:0004470 ... P40927 11790843
    15555 1gq2:K (2.5) BS01 NAP 1.1.1.40 GO:0004470 ... P40927 11790843
    15556 1gq2:K (2.5) BS02 OXL 1.1.1.40 GO:0004470 ... P40927 11790843
    15557 1gq2:K (2.5) BS03 MN 1.1.1.40 GO:0004470 ... P40927 11790843
    15558 1gq2:L (2.5) BS01 NAP 1.1.1.40 GO:0004470 ... P40927 11790843
    15559 1gq2:L (2.5) BS02 OXL 1.1.1.40 GO:0004470 ... P40927 11790843
    15560 1gq2:L (2.5) BS03 MN 1.1.1.40 GO:0004470 ... P40927 11790843
    15561 1gq2:M (2.5) BS01 NAP 1.1.1.40 GO:0004470 ... P40927 11790843
    15562 1gq2:M (2.5) BS02 OXL 1.1.1.40 GO:0004470 ... P40927 11790843
    15563 1gq2:M (2.5) BS03 MN 1.1.1.40 GO:0004470 ... P40927 11790843
    15564 1gq2:N (2.5) BS01 NAP 1.1.1.40 GO:0004470 ... P40927 11790843
    15565 1gq2:N (2.5) BS02 OXL 1.1.1.40 GO:0004470 ... P40927 11790843
    15566 1gq2:N (2.5) BS03 MN 1.1.1.40 GO:0004470 ... P40927 11790843
    15567 1gq2:O (2.5) BS01 NAP 1.1.1.40 GO:0004470 ... P40927 11790843
    15568 1gq2:O (2.5) BS02 OXL 1.1.1.40 GO:0004470 ... P40927 11790843
    15569 1gq2:O (2.5) BS03 MN 1.1.1.40 GO:0004470 ... P40927 11790843
    15570 1gq2:P (2.5) BS01 NAP 1.1.1.40 GO:0004470 ... P40927 11790843
    15571 1gq2:P (2.5) BS02 OXL 1.1.1.40 GO:0004470 ... P40927 11790843
    15572 1gq2:P (2.5) BS03 MN 1.1.1.40 GO:0004470 ... P40927 11790843
    15573 1gq6:A (1.75) BS01 MN 3.5.3.22 GO:0008783 ... P0DJQ3 12020346
    15574 1gq6:A (1.75) BS02 MN 3.5.3.22 GO:0008783 ... P0DJQ3 12020346
    15575 1gq6:B (1.75) BS01 MN 3.5.3.22 GO:0008783 ... P0DJQ3 12020346
    15576 1gq6:B (1.75) BS02 MN 3.5.3.22 GO:0008783 ... P0DJQ3 12020346
    15577 1gq6:C (1.75) BS01 MN 3.5.3.22 GO:0008783 ... P0DJQ3 12020346
    15578 1gq6:C (1.75) BS02 MN 3.5.3.22 GO:0008783 ... P0DJQ3 12020346
    15579 1gq7:A (2.45) BS01 MN 3.5.3.22 GO:0008783 ... P0DJQ3 12020346
    15580 1gq7:A (2.45) BS02 MN 3.5.3.22 GO:0008783 ... P0DJQ3 12020346
    15581 1gq7:B (2.45) BS01 MN 3.5.3.22 GO:0008783 ... P0DJQ3 12020346
    15582 1gq7:B (2.45) BS02 MN 3.5.3.22 GO:0008783 ... P0DJQ3 12020346
    15583 1gq7:C (2.45) BS01 MN 3.5.3.22 GO:0008783 ... P0DJQ3 12020346
    15584 1gq7:C (2.45) BS02 MN 3.5.3.22 GO:0008783 ... P0DJQ3 12020346
    15585 1gq7:D (2.45) BS01 MN 3.5.3.22 GO:0008783 ... P0DJQ3 12020346
    15586 1gq7:D (2.45) BS02 MN 3.5.3.22 GO:0008783 ... P0DJQ3 12020346
    15587 1gq7:E (2.45) BS01 MN 3.5.3.22 GO:0008783 ... P0DJQ3 12020346
    15588 1gq7:E (2.45) BS02 MN 3.5.3.22 GO:0008783 ... P0DJQ3 12020346
    15589 1gq7:F (2.45) BS01 MN 3.5.3.22 GO:0008783 ... P0DJQ3 12020346
    15590 1gq7:F (2.45) BS02 MN 3.5.3.22 GO:0008783 ... P0DJQ3 12020346
    15591 1gq9:A (2.6) BS01 MG 2.7.7.38 GO:0005737 ... P42216 11802716
    15592 1gq9:A (2.6) BS02 CTP 2.7.7.38 GO:0005737 ... P42216 11802716
    15593 1gq9:B (2.6) BS01 MG 2.7.7.38 GO:0005737 ... P42216 11802716
    15594 1gq9:B (2.6) BS02 CTP 2.7.7.38 GO:0005737 ... P42216 11802716
    15595 1gqa:A (1.8) BS01 HEC ? GO:0005506 ... P00148 N/A
    15596 1gqa:D (1.8) BS01 HEC ? GO:0005506 ... P00148 N/A
    15597 1gqb:A (1.8) BS01 BR ? GO:0005537 ... P11717 12499540
    15598 1gqb:A (1.8) BS02 BR ? GO:0005537 ... P11717 12499540
    15599 1gqb:B (1.8) BS01 BR ? GO:0005537 ... P11717 12499540
    15600 1gqb:B (1.8) BS02 BR ? GO:0005537 ... P11717 12499540

    Reference:
  • Chengxin Zhang, Xi Zhang, Peter L Freddolino, and Yang Zhang. BioLiP2: an updated structure database for biologically relevent ligand-protein interactions, Nucleic Acids Research, gkad630 (2023).
  • Jianyi Yang, Ambrish Roy, and Yang Zhang. BioLiP: a semi-manually curated database for biologically relevant ligand-protein interactions, Nucleic Acids Research, 41: D1096-D1103 (2013) (download the PDF file).
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