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●TM-score ●TM-align ●US-align ●MM-align ●RNA-align ●NW-align ●LS-align ●EDTSurf ●MVP ●MVP-Fit ●SPICKER ●HAAD ●PSSpred ●3DRobot ●MR-REX ●I-TASSER-MR ●SVMSEQ ●NeBcon ●ResPRE ●TripletRes ●DeepPotential ●WDL-RF ●ATPbind ●DockRMSD ●DeepMSA ●FASPR ●EM-Refiner ●GPU-I-TASSER

●BioLiP ●E. coli ●GLASS ●GPCR-HGmod ●GPCR-RD ●GPCR-EXP ●Tara-3D ●TM-fold ●DECOYS ●POTENTIAL ●RW/RWplus ●EvoEF ●HPSF ●THE-DB ●ADDRESS ●Alpaca-Antibody ●CASP7 ●CASP8 ●CASP9 ●CASP10 ●CASP11 ●CASP12 ●CASP13 ●CASP14

BioLiP
Download all results in tab-seperated text for 197122 receptor-ligand interactions, whose format is explained at readme.txt.
  • Hover over PDB to view the title of the structure. Click PDB to view the structure at the RCSB PDB database. Resolution -1.00 means the resolution is unavailable, e.g., for NMR structures.
  • Click Site # to view the binding site structure. Hover over Site # to view the binding residues.
  • Hover over Ligand to view the full ligand name. Click Ligand to view the 2D diagram and other detail information of the ligand.
  • Hover over EC number to view the full name of enzymatic activity.
  • Hover over GO terms to view all GO terms.
  • Hover over UniProt to view the protein name.

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    # PDB
    (Resolution Å)     		Site # Ligand EC number GO terms UniProt PubMed Binding
    affinity
    137201 6rz3:A (4.23) BS01 ZN ? GO:0000976 ... P04637 31395738
    137202 6rz4:A (2.7) BS02 NA ? GO:0004930 ... P0ABE7
    Q9Y271    		 31633023
    137203 6rz5:A (2.53) BS02 NA ? GO:0004930 ... P0ABE7
    Q9Y271    		 31633023
    137204 6rz5:B (2.53) BS03 NA ? GO:0004930 ... P0ABE7
    Q9Y271    		 31633023
    137205 6rza:A (4.5) BS02 MG 3.6.5.- GO:0000226 ... Q2XVP4 31264960
    137206 6rza:C (4.5) BS02 MG 3.6.5.- GO:0000226 ... Q2XVP4 31264960
    137207 6rzb:A (4.1) BS02 MG 3.6.5.- GO:0000226 ... Q2XVP4 31264960
    137208 6rzp:A (2.2) BS01 CA 3.4.21.64 GO:0004252 ... P06873 N/A
    137209 6rzr:A (1.9) BS01 ZN 3.5.2.6 GO:0008270 ... Q7WYA8 32205343
    137210 6rzr:A (1.9) BS02 ZN 3.5.2.6 GO:0008270 ... Q7WYA8 32205343
    137211 6rzr:B (1.9) BS01 ZN 3.5.2.6 GO:0008270 ... Q7WYA8 32205343
    137212 6rzr:B (1.9) BS02 ZN 3.5.2.6 GO:0008270 ... Q7WYA8 32205343
    137213 6rzs:A (2.2) BS02 ZN 3.5.2.6 GO:0008270 ... Q7WYA8 32205343
    137214 6rzs:A (2.2) BS03 ZN 3.5.2.6 GO:0008270 ... Q7WYA8 32205343
    137215 6rzs:B (2.2) BS03 ZN 3.5.2.6 GO:0008270 ... Q7WYA8 32205343
    137216 6rzs:B (2.2) BS04 ZN 3.5.2.6 GO:0008270 ... Q7WYA8 32205343
    137217 6rzx:A (1.0) BS01 ZN 4.2.1.1
    4.2.1.69    		 GO:0002009 ... P00918 32452709
    137218 6rzz:g (3.2) BS02 ZN ? GO:0002181 ... P87262 31115337
    137219 6rzz:i (3.2) BS03 ZN ? GO:0000448 ... P49166 31115337
    137220 6rzz:l (3.2) BS02 ZN ? GO:0002181 ... P0CX27 31115337
    137221 6rzz:m (3.2) BS02 ZN ? GO:0002181 ... P0CX25 31115337
    137222 6rzz:s (3.2) BS02 ZN 3.1.3.48 GO:0000027 ... Q02256 31115337
    137223 6rzz:u (3.2) BS02 ZN ? GO:0000278 ... P38344 31115337
    137224 6rzz:u (3.2) BS03 ZN ? GO:0000278 ... P38344 31115337
    137225 6rzz:w (3.2) BS02 ZN ? GO:0000055 ... P38861 31115337
    137226 6rzz:w (3.2) BS03 ZN ? GO:0000055 ... P38861 31115337
    137227 6s02:A (1.87) BS02 MG ? GO:0005524 ... K7NTP5 31690116
    137228 6s03:A (1.38) BS01 ZN 4.2.1.1
    4.2.1.69    		 GO:0002009 ... P00918 32452709
    137229 6s05:g (3.9) BS02 ZN ? GO:0002181 ... P87262 31115337
    137230 6s05:i (3.9) BS03 ZN ? GO:0000448 ... P49166 31115337
    137231 6s05:l (3.9) BS02 ZN ? GO:0002181 ... P0CX27 31115337
    137232 6s05:m (3.9) BS02 ZN ? GO:0002181 ... P0CX25 31115337
    137233 6s05:s (3.9) BS02 ZN 3.1.3.48 GO:0000027 ... Q02256 31115337
    137234 6s05:w (3.9) BS02 ZN ? GO:0000055 ... P38861 31115337
    137235 6s07:A (1.04) BS02 CU1 1.8.3.7 GO:0016491 ... D1A7C3 31588272
    137236 6s07:A (1.04) BS03 CA 1.8.3.7 GO:0016491 ... D1A7C3 31588272
    137237 6s07:A (1.04) BS04 CA 1.8.3.7 GO:0016491 ... D1A7C3 31588272
    137238 6s0d:A (1.52) BS01 MN 1.15.1.1 GO:0004784 ... P41977 N/A
    137239 6s0d:C (1.52) BS01 MN 1.15.1.1 GO:0004784 ... P41977 N/A
    137240 6s0h:A (2.85) BS01 ZN 3.5.2.6 GO:0008270 ... Q7WYA8 32205343
    137241 6s0h:A (2.85) BS02 ZN 3.5.2.6 GO:0008270 ... Q7WYA8 32205343
    137242 6s0h:B (2.85) BS01 ZN 3.5.2.6 GO:0008270 ... Q7WYA8 32205343
    137243 6s0h:B (2.85) BS02 ZN 3.5.2.6 GO:0008270 ... Q7WYA8 32205343
    137244 6s0i:A (1.803) BS01 ZN 3.1.2.6 GO:0004416 ... P0AC84 N/A
    137245 6s0i:A (1.803) BS02 ZN 3.1.2.6 GO:0004416 ... P0AC84 N/A
    137246 6s0k:f (3.1) BS02 ZN ? GO:0002181 ... P0A7Q6 31570874
    137247 6s0m:B (2.0) BS02 MG 3.1.13.- GO:0000287 ... P9WJ73 31299874
    137248 6s0m:B (2.0) BS03 MG 3.1.13.- GO:0000287 ... P9WJ73 31299874
    137249 6s0o:A (1.8) BS01 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137250 6s0o:A (1.8) BS02 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137251 6s0o:A (1.8) BS03 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137252 6s0o:B (1.8) BS01 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137253 6s0o:B (1.8) BS02 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137254 6s0o:B (1.8) BS03 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137255 6s0o:B (1.8) BS04 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137256 6s0o:B (1.8) BS05 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137257 6s0o:C (1.8) BS01 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137258 6s0o:C (1.8) BS02 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137259 6s0o:C (1.8) BS03 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137260 6s0o:D (1.8) BS01 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137261 6s0o:D (1.8) BS02 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137262 6s0o:D (1.8) BS03 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137263 6s0o:D (1.8) BS04 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137264 6s0o:E (1.8) BS01 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137265 6s0o:E (1.8) BS02 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137266 6s0o:E (1.8) BS03 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137267 6s0o:E (1.8) BS04 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137268 6s0o:E (1.8) BS05 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137269 6s0o:F (1.8) BS01 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137270 6s0o:F (1.8) BS02 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137271 6s0o:F (1.8) BS03 CU 1.10.3.2 GO:0005507 ... A0A0M4FJ81 31261802
    137272 6s0p:A (1.24) BS01 FE 1.13.11.20 GO:0005506 ... Q9SJI9 32868422
    137273 6s1x:A (1.76) BS01 ZN 3.4.17.21 GO:0004180 ... Q04609 32097010
    137274 6s1x:A (1.76) BS02 ZN 3.4.17.21 GO:0004180 ... Q04609 32097010
    137275 6s1x:A (1.76) BS03 CA 3.4.17.21 GO:0004180 ... Q04609 32097010
    137276 6s1y:A (2.2) BS01 ZN 3.4.-.- GO:0006508 ... A0NFU8 31682720
    137277 6s1z:A (2.5) BS02 ZN 3.4.-.- GO:0006508 ... A0NFU8 31682720
    137278 6s20:C (1.98) BS02 CA 3.1.6.- GO:0016787 ... Q8A2H2 32005816
    137279 6s21:A (2.8) BS04 CA 3.1.6.- GO:0004065 ... Q8A2F6 32005816
    137280 6s21:B (2.8) BS04 CA 3.1.6.- GO:0004065 ... Q8A2F6 32005816
    137281 6s21:C (2.8) BS03 CA 3.1.6.- GO:0004065 ... Q8A2F6 32005816
    137282 6s22:A (1.96) BS03 MN 2.4.1.41 N/A H0ZAB5 31932717
    137283 6s23:B (2.38) BS03 MG ? GO:0000166 ... M2XAQ9 31930452
    137284 6s24:A (2.12) BS03 MN 2.4.1.41 N/A H0ZAB5 31932717
    137285 6s2t:A (2.75) BS02 MN 2.7.6.5 GO:0015969 ... Q5SHL3 32393900
    137286 6s2t:A (2.75) BS03 MG 2.7.6.5 GO:0015969 ... Q5SHL3 32393900
    137287 6s2u:A (2.95) BS01 MN 2.7.6.5 GO:0015969 ... Q5SHL3 32393900
    137288 6s2v:A (2.96) BS01 MN 2.7.6.5 GO:0015969 ... Q5SHL3 32393900
    137289 6s2v:B (2.96) BS01 MN 2.7.6.5 GO:0015969 ... Q5SHL3 32393900
    137290 6s2v:C (2.96) BS01 MN 2.7.6.5 GO:0015969 ... Q5SHL3 32393900
    137291 6s39:A (1.88) BS03 CA ? GO:0000122 ... P31947 32501690
    137292 6s3a:A (1.52) BS02 ZN 2.7.7.48
    3.4.22.28
    3.4.22.29
    3.6.1.15    		 GO:0003723 ... P03313 34985963
    137293 6s3g:A (1.9) BS01 ZN 3.1.1.3 GO:0004806 ... Q93A71 N/A
    137294 6s3g:A (1.9) BS02 CA 3.1.1.3 GO:0004806 ... Q93A71 N/A
    137295 6s3j:A (1.9) BS01 ZN 3.1.1.3 GO:0004806 ... Q93A71 N/A
    137296 6s3j:A (1.9) BS02 CA 3.1.1.3 GO:0004806 ... Q93A71 N/A
    137297 6s3k:A (3.7) BS01 K ? GO:0005886 ... P96589 32005818
    137298 6s3k:A (3.7) BS02 K ? GO:0005886 ... P96589 32005818
    137299 6s3k:A (3.7) BS03 K ? GO:0005886 ... P96589 32005818
    137300 6s3k:B (3.7) BS01 K ? GO:0005886 ... P96589 32005818
    137301 6s3k:B (3.7) BS02 K ? GO:0005886 ... P96589 32005818
    137302 6s3k:B (3.7) BS03 K ? GO:0005886 ... P96589 32005818
    137303 6s3n:B (2.533) BS03 MG ? GO:0000723 ... K7RJ88 33784404
    137304 6s3v:A (2.0) BS01 ZN 3.1.1.3 GO:0004806 ... Q93A71 N/A
    137305 6s3v:A (2.0) BS02 CA 3.1.1.3 GO:0004806 ... Q93A71 N/A
    137306 6s40:A (1.9) BS02 MG ? GO:0000122 ... P31947 32501690
    137307 6s40:A (1.9) BS06 CA ? GO:0000122 ... P31947 32501690
    137308 6s48:A (1.9) BS08 CA ? GO:0003677 ... Q8YYB7 32459314
    137309 6s48:A (1.9) BS09 CA ? GO:0003677 ... Q8YYB7 32459314
    137310 6s48:B (1.9) BS06 CA ? GO:0003677 ... Q8YYB7 32459314
    137311 6s4b:A (1.6) BS02 CA ? N/A O60885 33275431
    137312 6s4c:A (2.0) BS01 MG ? N/A Q63870 32031736
    137313 6s53:A (2.8) BS01 ZN 2.3.2.27 GO:0004842 ... P19474 31582740
    137314 6s53:A (2.8) BS02 ZN 2.3.2.27 GO:0004842 ... P19474 31582740
    137315 6s53:B (2.8) BS01 ZN 2.3.2.27 GO:0004842 ... P19474 31582740
    137316 6s53:B (2.8) BS02 ZN 2.3.2.27 GO:0004842 ... P19474 31582740
    137317 6s53:G (2.8) BS01 ZN 2.3.2.27 GO:0004842 ... P19474 31582740
    137318 6s53:G (2.8) BS02 ZN 2.3.2.27 GO:0004842 ... P19474 31582740
    137319 6s53:H (2.8) BS01 ZN 2.3.2.27 GO:0004842 ... P19474 31582740
    137320 6s53:H (2.8) BS02 ZN 2.3.2.27 GO:0004842 ... P19474 31582740
    137321 6s58:D (2.32) BS01 CA ? GO:0003677 ... Q8YYB7 32459314
    137322 6s58:D (2.32) BS02 CA ? GO:0003677 ... Q8YYB7 32459314
    137323 6s5c:A (3.0) BS02 CA ? GO:0005515 ... O32080 31868587
    137324 6s5c:B (3.0) BS01 CA ? GO:0005515 ... O32080 31868587
    137325 6s5f:A (1.7) BS03 MG ? GO:0003924 ... Q96DA2 N/A
    137326 6s5h:A (2.0) BS02 MG ? GO:0003924 ... P57729 N/A
    137327 6s5p:A (1.46) BS01 CA ? GO:0030246 ... Q9HYN5 30830745
    137328 6s5p:A (1.46) BS02 CA ? GO:0030246 ... Q9HYN5 30830745
    137329 6s5p:B (1.46) BS01 CA ? GO:0030246 ... Q9HYN5 30830745
    137330 6s5p:B (1.46) BS02 CA ? GO:0030246 ... Q9HYN5 30830745
    137331 6s5p:C (1.46) BS01 CA ? GO:0030246 ... Q9HYN5 30830745
    137332 6s5p:C (1.46) BS02 CA ? GO:0030246 ... Q9HYN5 30830745
    137333 6s5p:D (1.46) BS01 CA ? GO:0030246 ... Q9HYN5 30830745
    137334 6s5p:D (1.46) BS03 CA ? GO:0030246 ... Q9HYN5 30830745
    137335 6s5r:A (2.076) BS01 CA ? GO:0030246 ... Q9HYN5 N/A
    137336 6s5r:A (2.076) BS02 CA ? GO:0030246 ... Q9HYN5 N/A
    137337 6s5r:B (2.076) BS01 CA ? GO:0030246 ... Q9HYN5 N/A
    137338 6s5r:B (2.076) BS03 CA ? GO:0030246 ... Q9HYN5 N/A
    137339 6s5r:C (2.076) BS01 CA ? GO:0030246 ... Q9HYN5 N/A
    137340 6s5r:C (2.076) BS02 CA ? GO:0030246 ... Q9HYN5 N/A
    137341 6s5r:D (2.076) BS01 CA ? GO:0030246 ... Q9HYN5 N/A
    137342 6s5r:D (2.076) BS02 CA ? GO:0030246 ... Q9HYN5 N/A
    137343 6s5s:A (1.433) BS01 CA ? GO:0030246 ... Q9HYN5 N/A
    137344 6s5s:A (1.433) BS02 CA ? GO:0030246 ... Q9HYN5 N/A
    137345 6s61:A (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137346 6s61:B (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137347 6s61:C (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137348 6s61:D (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137349 6s61:E (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137350 6s61:F (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137351 6s61:G (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137352 6s61:H (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137353 6s61:I (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137354 6s61:J (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137355 6s61:K (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137356 6s61:L (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137357 6s61:M (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137358 6s61:N (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137359 6s61:O (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137360 6s61:P (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137361 6s61:Q (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137362 6s61:R (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137363 6s61:S (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137364 6s61:T (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137365 6s61:U (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137366 6s61:V (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137367 6s61:W (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137368 6s61:X (1.84) BS01 ZN 1.16.3.1 GO:0004322 ... P09528 N/A
    137369 6s63:A (1.85) BS02 CA ? GO:0005216 ... P96787 34981761
    137370 6s6a:A (2.63) BS02 MG 3.6.5.- GO:0003924 ... Q7L523 31601764
    137371 6s6a:B (2.63) BS02 MG 3.6.5.- GO:0003924 ... Q7L523 31601764
    137372 6s6b:K (2.75) BS01 ZN ? GO:0016020 ... F0NDX2 32730741
    137373 6s6c:A (1.07) BS02 CA ? GO:0005216 ... P96787 33504778
    137374 6s6d:A (2.5) BS02 MG 3.6.5.- GO:0003924 ... Q7L523 31601764
    137375 6s6d:B (2.5) BS02 MG 3.6.5.- GO:0003924 ... Q7L523 31601764
    137376 6s6v:A (3.5) BS01 MN ? GO:0000014 ... P0AG76 31492634
    137377 6s6v:A (3.5) BS02 MN ? GO:0000014 ... P0AG76 31492634
    137378 6s6v:B (3.5) BS01 MN ? GO:0000014 ... P0AG76 31492634
    137379 6s6v:B (3.5) BS02 MN ? GO:0000014 ... P0AG76 31492634
    137380 6s6v:C (3.5) BS02 MG ? GO:0000014 ... P13458 31492634
    137381 6s6v:D (3.5) BS03 MG ? GO:0000014 ... P13458 31492634
    137382 6s6y:A (3.1) BS02 ZN N/A GO:0016810 ... N/A 31776258
    137383 6s6y:A (3.1) BS03 ZN N/A GO:0016810 ... N/A 31776258
    137384 6s6y:A (3.1) BS04 CA N/A GO:0016810 ... N/A 31776258
    137385 6s6y:A (3.1) BS05 CA N/A GO:0016810 ... N/A 31776258
    137386 6s6y:B (3.1) BS01 CA N/A N/A N/A 31776258
    137387 6s6y:E (3.1) BS01 ZN N/A GO:0016810 ... N/A 31776258
    137388 6s6y:E (3.1) BS02 ZN N/A GO:0016810 ... N/A 31776258
    137389 6s6y:I (3.1) BS02 ZN N/A GO:0016810 ... N/A 31776258
    137390 6s6y:I (3.1) BS03 ZN N/A GO:0016810 ... N/A 31776258
    137391 6s6y:L (3.1) BS01 CA N/A GO:0006730 ... N/A 31776258
    137392 6s6y:M (3.1) BS01 ZN N/A GO:0016810 ... N/A 31776258
    137393 6s6z:A (2.0) BS01 MG 3.2.1.23 GO:0003824 ... Q56307 31874027
    137394 6s6z:B (2.0) BS01 MG 3.2.1.23 GO:0003824 ... Q56307 31874027
    137395 6s6z:C (2.0) BS01 MG 3.2.1.23 GO:0003824 ... Q56307 31874027
    137396 6s6z:D (2.0) BS01 MG 3.2.1.23 GO:0003824 ... Q56307 31874027
    137397 6s6z:E (2.0) BS01 MG 3.2.1.23 GO:0003824 ... Q56307 31874027
    137398 6s6z:F (2.0) BS01 MG 3.2.1.23 GO:0003824 ... Q56307 31874027
    137399 6s6z:G (2.0) BS01 MG 3.2.1.23 GO:0003824 ... Q56307 31874027
    137400 6s6z:H (2.0) BS01 MG 3.2.1.23 GO:0003824 ... Q56307 31874027
    Reference:
  • Chengxin Zhang, Xi Zhang, Peter L Freddolino, and Yang Zhang. BioLiP2: an updated structure database for biologically relevent ligand-protein interactions, Nucleic Acids Research, gkad630 (2023).
  • Jianyi Yang, Ambrish Roy, and Yang Zhang. BioLiP: a semi-manually curated database for biologically relevant ligand-protein interactions, Nucleic Acids Research, 41: D1096-D1103 (2013) (download the PDF file).

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