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BioLiP
Download all results in tab-seperated text for 287 receptor-ligand interactions, whose format is explained at readme.txt.
  • Hover over PDB to view the title of the structure. Click PDB to view the structure at the RCSB PDB database. Resolution -1.00 means the resolution is unavailable, e.g., for NMR structures.
  • Click Site # to view the binding site structure. Hover over Site # to view the binding residues.
  • Hover over Ligand to view the full ligand name. Click Ligand to view the 2D diagram and other detail information of the ligand.
  • Hover over EC number to view the full name of enzymatic activity.
  • Hover over GO terms to view all GO terms.
  • Hover over UniProt to view the protein name.
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    # PDB
    (Resolution Å)
    Site # Ligand EC number GO terms UniProt PubMed Binding
    affinity
    1 1a9y:A (1.8) BS02 UPG 5.1.3.2 GO:0003978 ... P09147 9708982
    2 1a9z:A (1.9) BS02 UPG 5.1.3.2 GO:0003978 ... P09147 9708982
    3 1ek6:A (1.5) BS02 UPG 5.1.3.2
    5.1.3.7
    GO:0003974 ... Q14376 10801319
    4 1ek6:B (1.5) BS02 UPG 5.1.3.2
    5.1.3.7
    GO:0003974 ... Q14376 10801319
    5 1g93:A (2.5) BS03 UPG 2.4.1.87 GO:0005975 ... P14769 11179209
    6 1guq:A (1.8) BS04 UPG 2.7.7.12 GO:0004335 ... P09148 9063869
    7 1guq:A (1.8) BS05 UPG 2.7.7.12 GO:0004335 ... P09148 9063869
    8 1guq:B (1.8) BS01 UPG 2.7.7.12 GO:0004335 ... P09148 9063869
    9 1guq:B (1.8) BS05 UPG 2.7.7.12 GO:0004335 ... P09148 9063869
    10 1guq:C (1.8) BS04 UPG 2.7.7.12 GO:0004335 ... P09148 9063869
    11 1guq:C (1.8) BS05 UPG 2.7.7.12 GO:0004335 ... P09148 9063869
    12 1guq:D (1.8) BS01 UPG 2.7.7.12 GO:0004335 ... P09148 9063869
    13 1guq:D (1.8) BS05 UPG 2.7.7.12 GO:0004335 ... P09148 9063869
    14 1i24:A (1.2) BS02 UPG 3.13.1.1 N/A O48917 N/A
    15 1i2b:A (1.75) BS02 UPG 3.13.1.1 N/A O48917 N/A
    16 1i2c:A (1.6) BS02 UPG 3.13.1.1 N/A O48917 N/A
    17 1i3k:A (1.5) BS02 UPG 5.1.3.2
    5.1.3.7
    GO:0003974 ... Q14376 11279193
    18 1i3k:B (1.5) BS02 UPG 5.1.3.2
    5.1.3.7
    GO:0003974 ... Q14376 11279193
    19 1iin:A (2.1) BS01 UPG 2.7.7.24 GO:0000287 ... P26393 11373625
    20 1iin:B (2.1) BS01 UPG 2.7.7.24 GO:0000287 ... P26393 11373625
    21 1iin:C (2.1) BS01 UPG 2.7.7.24 GO:0000287 ... P26393 11373625
    22 1iin:D (2.1) BS01 UPG 2.7.7.24 GO:0000287 ... P26393 11373625
    23 1j39:A (1.87) BS01 UPG 2.4.1.27 GO:0006304 ... P04547 12860129
    24 1kvq:A (2.15) BS02 UPG 5.1.3.2 GO:0003978 ... P09147 9271499
    25 1kvs:A (2.15) BS02 UPG 5.1.3.2 GO:0003978 ... P09147 9271499
    26 1kvt:A (2.15) BS02 UPG 5.1.3.2 GO:0003978 ... P09147 9271499
    27 1kvu:A (1.9) BS02 UPG 5.1.3.2 GO:0003978 ... P09147 9271498
    28 1ll2:A (1.9) BS02 UPG 2.4.1.186 GO:0005737 ... P13280 12051921
    29 1lrl:A (1.8) BS02 UPG 5.1.3.2 GO:0003978 ... P09147 12019271
    30 1mp4:A (2.2) BS01 UPG 2.7.7.24 GO:0000166 ... Q9F7G8 12374866
    31 1mp4:B (2.2) BS01 UPG 2.7.7.24 GO:0000166 ... Q9F7G8 12374866
    32 1mp5:A (2.75) BS01 UPG 2.7.7.24 GO:0000287 ... P26393 12374866
    33 1mp5:B (2.75) BS01 UPG 2.7.7.24 GO:0000287 ... P26393 12374866
    34 1mp5:C (2.75) BS01 UPG 2.7.7.24 GO:0000287 ... P26393 12374866
    35 1mp5:D (2.75) BS01 UPG 2.7.7.24 GO:0000287 ... P26393 12374866
    36 1nzd:A (2.0) BS01 UPG 2.4.1.27 GO:0006304 ... P04547 12860129
    37 1nzf:A (2.1) BS01 UPG 2.4.1.27 GO:0006304 ... P04547 12860129
    38 1o23:B (2.32) BS02 UPG 2.4.1.-
    2.4.1.22
    2.4.1.275
    2.4.1.38
    2.4.1.90
    GO:0005975 ... P08037 11485999
    39 1o23:D (2.32) BS02 UPG 2.4.1.-
    2.4.1.22
    2.4.1.275
    2.4.1.38
    2.4.1.90
    GO:0005975 ... P08037 11485999
    40 1qrr:A (1.6) BS02 UPG 3.13.1.1 N/A O48917 10557279
    41 1sb9:A (2.5) BS02 UPG 5.1.3.7 GO:0000166 ... Q8KN66 15016816
    42 1uqu:A (2.0) BS01 UPG 2.4.1.15 GO:0003824 ... P31677 14570926
    43 1xel:A (1.8) BS02 UPG 5.1.3.2 GO:0003978 ... P09147 8611497
    44 1y6f:A (2.4) BS03 UPG 2.4.1.26 GO:0006304 ... P04519 16081100
    45 1z45:A (1.85) BS03 UPG 5.1.3.2
    5.1.3.3
    GO:0003824 ... P04397 15795221
    46 1zdf:A (2.45) BS02 UPG 2.4.1.186 GO:0005737 ... P13280 15849187
    47 1zdg:A (2.3) BS02 UPG 2.4.1.186 GO:0005737 ... P13280 15849187
    48 2acw:A (2.6) BS01 UPG 2.4.1.- GO:0000166 ... Q5IFH7 16214900
    49 2acw:B (2.6) BS01 UPG 2.4.1.- GO:0000166 ... Q5IFH7 16214900
    50 2am3:A (1.8) BS02 UPG 2.4.1.101 GO:0006486 ... P27115 16769084
    51 2gn9:A (2.8) BS02 UPG 4.2.1.115 GO:0000166 ... O25511 16651261
    52 2gn9:B (2.8) BS02 UPG 4.2.1.115 GO:0000166 ... O25511 16651261
    53 2icy:A (1.64) BS01 UPG 2.7.7.9 GO:0003983 ... Q9M9P3 17178129
    54 2icy:B (1.64) BS01 UPG 2.7.7.9 GO:0003983 ... Q9M9P3 17178129
    55 2oeg:A (2.3) BS01 UPG 2.7.7.9 GO:0003983 ... Q4QDU3 17303565
    56 2p72:B (2.0) BS02 UPG ? GO:0016020 ... Q89399 17850743
    57 2pa4:A (2.0) BS01 UPG N/A GO:0003983 ... N/A 17567737
    58 2pa4:B (2.0) BS01 UPG N/A GO:0003983 ... N/A 17567737
    59 2pa4:C (2.0) BS01 UPG N/A GO:0003983 ... N/A 17567737
    60 2pa4:D (2.0) BS01 UPG N/A GO:0003983 ... N/A 17567737
    61 2q3e:A (2.0) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21502315
    62 2q3e:B (2.0) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21502315
    63 2q3e:C (2.0) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21502315
    64 2q3e:D (2.0) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21502315
    65 2q3e:E (2.0) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21502315
    66 2q3e:F (2.0) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21502315
    67 2q3e:G (2.0) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21502315
    68 2q3e:H (2.0) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21502315
    69 2q3e:I (2.0) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21502315
    70 2q3e:J (2.0) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21502315
    71 2q3e:K (2.0) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21502315
    72 2q3e:L (2.0) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21502315
    73 2vkd:A (2.53) BS02 UPG 2.4.1.-
    3.4.22.-
    GO:0016757 ... Q46342 18325534
    74 2vkd:B (2.53) BS02 UPG 2.4.1.-
    3.4.22.-
    GO:0016757 ... Q46342 18325534
    75 2vkd:C (2.53) BS02 UPG 2.4.1.-
    3.4.22.-
    GO:0016757 ... Q46342 18325534
    76 2vkh:A (2.3) BS02 UPG 2.4.1.-
    3.4.22.-
    GO:0016757 ... Q46342 18325534
    77 2vkh:B (2.3) BS02 UPG 2.4.1.-
    3.4.22.-
    GO:0016757 ... Q46342 18325534
    78 2vkh:C (2.3) BS02 UPG 2.4.1.-
    3.4.22.-
    GO:0016757 ... Q46342 18325534
    79 2wzg:A (1.9) BS01 UPG ? GO:0000166 ... Q5WWY0 20030628 MOAD: Kd=100nM
    80 2xa2:A (2.5) BS01 UPG 2.4.1.245 GO:0006006 ... O58762 20888836
    81 2xa2:B (2.5) BS01 UPG 2.4.1.245 GO:0006006 ... O58762 20888836
    82 2xa9:A (2.5) BS01 UPG 2.4.1.245 GO:0006006 ... O58762 20888836
    83 2xa9:B (2.5) BS01 UPG 2.4.1.245 GO:0006006 ... O58762 20888836
    84 3ckq:A (3.0) BS02 UPG 2.4.1.266 GO:0016757 ... Q73WU1 18667419
    85 3enk:A (1.9) BS02 UPG 5.1.3.2 GO:0000166 ... Q3JPI3 N/A
    86 3enk:B (1.9) BS02 UPG 5.1.3.2 GO:0000166 ... Q3JPI3 N/A
    87 3gf4:A (2.45) BS02 UPG 5.4.99.9 GO:0008767 ... Q48485 19500588 MOAD: Kd=750uM
    88 3gue:A (1.92) BS01 UPG 2.7.7.9 GO:0003983 ... Q388T4 N/A
    89 3gue:B (1.92) BS01 UPG 2.7.7.9 GO:0003983 ... Q388T4 N/A
    90 3jsz:A (1.7) BS01 UPG ? GO:0000166 ... Q5WWY0 19941871
    91 3juk:A (2.3) BS01 UPG 2.7.7.9 GO:0003983 ... O25363 20238176 PDBbind: -logKd/Ki=4.89, Kd=12.9uM
    92 3juk:B (2.3) BS01 UPG 2.7.7.9 GO:0003983 ... O25363 20238176
    93 3juk:C (2.3) BS01 UPG 2.7.7.9 GO:0003983 ... O25363 20238176
    94 3juk:D (2.3) BS01 UPG 2.7.7.9 GO:0003983 ... O25363 20238176
    95 3khu:A (2.3) BS01 UPG 1.1.1.22 GO:0001702 ... O60701 22123821
    96 3khu:A (2.3) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 22123821
    97 3khu:B (2.3) BS01 UPG 1.1.1.22 GO:0001702 ... O60701 22123821
    98 3khu:B (2.3) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 22123821
    99 3khu:C (2.3) BS01 UPG 1.1.1.22 GO:0001702 ... O60701 22123821
    100 3khu:C (2.3) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 22123821
    101 3khu:D (2.3) BS01 UPG 1.1.1.22 GO:0001702 ... O60701 22123821
    102 3khu:D (2.3) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 22123821
    103 3khu:E (2.3) BS01 UPG 1.1.1.22 GO:0001702 ... O60701 22123821
    104 3khu:E (2.3) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 22123821
    105 3khu:F (2.3) BS01 UPG 1.1.1.22 GO:0001702 ... O60701 22123821
    106 3khu:F (2.3) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 22123821
    107 3oh4:A (2.21) BS01 UPG 2.7.7.64 GO:0016740 ... D3G6S4 21073876
    108 3rmw:A (1.93) BS01 UPG 2.4.1.186 GO:0016757 ... P46976 22160680
    109 3srz:A (2.58) BS02 UPG 2.4.1.-
    3.4.22.-
    GO:0016757 ... P16154 22267739
    110 3t7o:A (1.85) BS03 UPG 2.4.1.186 GO:0016757 ... P46976 22160680
    111 3t7o:B (1.85) BS02 UPG 2.4.1.186 GO:0016757 ... P46976 22160680
    112 3tdk:A (2.8) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21984906
    113 3tdk:B (2.8) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21984906
    114 3tdk:C (2.8) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21984906
    115 3tdk:D (2.8) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21984906
    116 3tdk:E (2.8) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21984906
    117 3tdk:F (2.8) BS01 UPG 1.1.1.22 GO:0001702 ... O60701 21984906
    118 3tdk:G (2.8) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21984906
    119 3tdk:H (2.8) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21984906
    120 3tdk:I (2.8) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21984906
    121 3tdk:J (2.8) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21984906
    122 3tdk:K (2.8) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21984906
    123 3tdk:L (2.8) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 21984906
    124 3v91:A (2.0) BS02 UPG 2.4.1.186 GO:0016757 ... P13280 22226635
    125 3vtf:A (2.0) BS01 UPG 1.1.1.22 GO:0000271 ... A1RUM9 22949183
    126 4e7b:A (2.0) BS01 UPG 2.5.1.7 GO:0003824 ... P33038 N/A
    127 4e7b:B (2.0) BS01 UPG 2.5.1.7 GO:0003824 ... P33038 N/A
    128 4e7b:C (2.0) BS01 UPG 2.5.1.7 GO:0003824 ... P33038 N/A
    129 4e7b:D (2.0) BS01 UPG 2.5.1.7 GO:0003824 ... P33038 N/A
    130 4e7e:A (2.3) BS01 UPG 2.5.1.7 GO:0003824 ... P33038 N/A
    131 4e7e:D (2.3) BS01 UPG 2.5.1.7 GO:0003824 ... P33038 N/A
    132 4edf:A (2.08) BS01 UPG 1.1.1.22 GO:0001702 ... O60701 23106432 MOAD: Kd=33uM
    133 4edf:B (2.08) BS01 UPG 1.1.1.22 GO:0001702 ... O60701 23106432 MOAD: Kd=33uM
    134 4edf:C (2.08) BS01 UPG 1.1.1.22 GO:0001702 ... O60701 23106432 MOAD: Kd=33uM
    135 4edf:D (2.08) BS01 UPG 1.1.1.22 GO:0001702 ... O60701 23106432 MOAD: Kd=33uM
    136 4ho6:A (1.92) BS01 UPG 2.7.7.24 GO:0000166 ... Q9AGY4 N/A
    137 4ho8:A (2.6) BS01 UPG 2.7.7.24 GO:0000166 ... Q9AGY4 N/A
    138 4ho8:B (2.6) BS01 UPG 2.7.7.24 GO:0000166 ... Q9AGY4 N/A
    139 4ho8:C (2.6) BS01 UPG 2.7.7.24 GO:0000166 ... Q9AGY4 N/A
    140 4ho8:D (2.6) BS01 UPG 2.7.7.24 GO:0000166 ... Q9AGY4 N/A
    141 4lis:A (2.8) BS01 UPG 5.1.3.2 GO:0000166 ... C8VAU8 24116166
    142 4m2a:A (1.66) BS01 UPG 2.7.7.9 GO:0003983 ... Q4QDU3 N/A
    143 4m2b:A (2.2) BS01 UPG 2.7.7.9 GO:0003983 ... Q4QDU3 N/A
    144 4r7p:A (3.35) BS01 UPG 2.7.7.9 GO:0003983 ... Q16851 25860585
    145 4wma:A (1.62) BS04 UPG 2.4.2.62 GO:0016757 ... Q3U4G3 26414444
    146 4xsr:A (2.39) BS01 UPG ? GO:0000166 ... Q8YQW3 26692049
    147 4xsr:B (2.39) BS01 UPG ? GO:0000166 ... Q8YQW3 26692049
    148 4y6n:A (2.348) BS02 UPG 2.4.1.266 GO:0000287 ... P9WMW9 26136334
    149 4y6u:A (2.271) BS02 UPG 2.4.1.266 GO:0000287 ... P9WMW9 26136334
    150 4y7f:A (3.231) BS03 UPG 2.4.1.266 GO:0000287 ... P9WMW9 26136334
    151 4y7g:A (2.59) BS03 UPG 2.4.1.266 GO:0000287 ... P9WMW9 26136334
    152 4y9x:A (2.637) BS02 UPG 2.4.1.266 GO:0000287 ... P9WMW9 26136334
    153 4zrn:A (2.0) BS02 UPG 5.1.3.2 GO:0000166 ... Q9WYX9 26344854
    154 4zrn:B (2.0) BS02 UPG 5.1.3.2 GO:0000166 ... Q9WYX9 26344854
    155 5c4b:A (1.54) BS02 UPG 2.4.1.37
    2.4.1.40
    GO:0005975 ... P16442 26374898
    156 5c4c:A (1.43) BS01 UPG 2.4.1.37
    2.4.1.40
    GO:0005975 ... P16442 26374898
    157 5c4d:A (1.4) BS03 UPG 2.4.1.37
    2.4.1.40
    GO:0005975 ... P16442 26374898
    158 5c4f:A (1.41) BS03 UPG 2.4.1.37
    2.4.1.40
    GO:0005975 ... P16442 26374898
    159 5c8r:A (1.45) BS03 UPG 2.4.1.37
    2.4.1.40
    GO:0005975 ... P16442 26374898
    160 5gl5:A (1.9) BS01 UPG 2.4.1.-
    2.4.1.173
    GO:0005975 ... Q06321 30395931
    161 5gl5:B (1.9) BS01 UPG 2.4.1.-
    2.4.1.173
    GO:0005975 ... Q06321 30395931
    162 5h18:A (1.4) BS01 UPG ? GO:0000166 ... A0A2D0TCJ6 28939828
    163 5hut:A (1.9) BS01 UPG 2.4.1.15 GO:0003824 ... Q92410 28743811
    164 5hut:B (1.9) BS01 UPG 2.4.1.15 GO:0003824 ... Q92410 28743811
    165 5huv:A (2.002) BS01 UPG 2.4.1.15 GO:0003824 ... Q92410 N/A
    166 5huv:B (2.002) BS01 UPG 2.4.1.15 GO:0003824 ... Q92410 N/A
    167 5i1f:A (2.15) BS01 UPG 2.7.7.9 GO:0003983 ... A4JT02 N/A
    168 5jsx:A (2.81) BS02 UPG 2.4.1.266 GO:0016757 ... A5U1Q6 28625787
    169 5lor:A (2.192) BS01 UPG 3.6.1.45 GO:0005515 ... Q9BRQ3 N/A
    170 5nzg:A (1.6) BS01 UPG 2.7.7.9 GO:0003983 ... Q4QDU3 N/A
    171 5nzi:A (2.05) BS01 UPG 2.7.7.9 GO:0003983 ... Q4QDU3 N/A
    172 5nzj:A (1.95) BS01 UPG 2.7.7.9 GO:0003983 ... Q4QDU3 N/A
    173 5tjh:C (2.05) BS01 UPG 1.1.1.22 GO:0001702 ... O60701 27966912
    174 6c4j:A (2.53) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 30457329
    175 6c4j:B (2.53) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 30457329
    176 6c4j:C (2.53) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 30457329
    177 6c4j:D (2.53) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 30457329
    178 6c4j:E (2.53) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 30457329
    179 6c4j:F (2.53) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 30457329
    180 6c4j:G (2.53) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 30457329
    181 6c4j:H (2.53) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 30457329
    182 6c4j:I (2.53) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 30457329
    183 6c4j:J (2.53) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 30457329
    184 6c4j:K (2.53) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 30457329
    185 6c4j:L (2.53) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 30457329
    186 6c5a:A (2.3) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 N/A
    187 6c5a:B (2.3) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 N/A
    188 6c5a:C (2.3) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 N/A
    189 6c5a:D (2.3) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 N/A
    190 6c5a:E (2.3) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 N/A
    191 6c5a:F (2.3) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 N/A
    192 6c5z:A (2.95) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 N/A
    193 6c5z:B (2.95) BS01 UPG 1.1.1.22 GO:0001702 ... O60701 N/A
    194 6c5z:C (2.95) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 N/A
    195 6c5z:D (2.95) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 N/A
    196 6c5z:E (2.95) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 N/A
    197 6c5z:F (2.95) BS02 UPG 1.1.1.22 GO:0001702 ... O60701 N/A
    198 6d9b:A (2.2) BS02 UPG N/A GO:0008767 ... N/A N/A
    199 6d9c:A (3.05) BS02 UPG N/A GO:0008767 ... N/A N/A
    200 6fsn:A (1.19) BS02 UPG ? GO:0003980 ... G0SB58 N/A

    Reference:
  • Chengxin Zhang, Xi Zhang, Peter L Freddolino, and Yang Zhang. BioLiP2: an updated structure database for biologically relevent ligand-protein interactions, Nucleic Acids Research, gkad630 (2023).
  • Jianyi Yang, Ambrish Roy, and Yang Zhang. BioLiP: a semi-manually curated database for biologically relevant ligand-protein interactions, Nucleic Acids Research, 41: D1096-D1103 (2013) (download the PDF file).
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