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BioLiP
Download all results in tab-seperated text for 3495 receptor-ligand interactions, whose format is explained at readme.txt.
  • Hover over PDB to view the title of the structure. Click PDB to view the structure at the RCSB PDB database. Resolution -1.00 means the resolution is unavailable, e.g., for NMR structures.
  • Click Site # to view the binding site structure. Hover over Site # to view the binding residues.
  • Hover over Ligand to view the full ligand name. Click Ligand to view the 2D diagram and other detail information of the ligand.
  • Hover over EC number to view the full name of enzymatic activity.
  • Hover over GO terms to view all GO terms.
  • Hover over UniProt to view the protein name.

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    # PDB
    (Resolution Å)     		Site # Ligand EC number GO terms UniProt PubMed Binding
    affinity
    1601 3zpc:A (2.2) BS02 PO4 1.1.1.193
    3.5.4.26    		 GO:0003824 ... D0CB74 23722836
    1602 3zpc:A (2.2) BS03 PO4 1.1.1.193
    3.5.4.26    		 GO:0003824 ... D0CB74 23722836
    1603 3zpc:B (2.2) BS02 PO4 1.1.1.193
    3.5.4.26    		 GO:0003824 ... D0CB74 23722836
    1604 3zpc:B (2.2) BS03 PO4 1.1.1.193
    3.5.4.26    		 GO:0003824 ... D0CB74 23722836
    1605 3zs7:A (2.0) BS01 PO4 2.7.1.35 GO:0005524 ... O15927 22857512
    1606 3zu0:A (2.001) BS03 PO4 3.1.3.5 GO:0000166 ... P44569 21933152
    1607 3zzm:A (2.2) BS01 PO4 2.1.2.3
    3.5.4.10    		 GO:0003824 ... P9WHM7 21956117
    1608 4a1o:A (2.48) BS02 PO4 2.1.2.3
    3.5.4.10    		 GO:0003824 ... P9WHM7 21956117
    1609 4a1o:B (2.48) BS02 PO4 2.1.2.3
    3.5.4.10    		 GO:0003824 ... P9WHM7 21956117
    1610 4a1o:B (2.48) BS04 PO4 2.1.2.3
    3.5.4.10    		 GO:0003824 ... P9WHM7 21956117
    1611 4a6f:A (1.68) BS01 PO4 ? N/A P40485 22574179
    1612 4a6f:A (1.68) BS03 PO4 ? N/A P40485 22574179
    1613 4a6h:A (1.449) BS01 PO4 ? N/A P40485 22574179
    1614 4a6h:B (1.449) BS01 PO4 ? N/A P40485 22574179
    1615 4a6h:B (1.449) BS02 PO4 ? N/A P40485 22574179
    1616 4a6h:C (1.449) BS01 PO4 ? N/A P40485 22574179
    1617 4a6h:C (1.449) BS02 PO4 ? N/A P40485 22574179
    1618 4a6h:D (1.449) BS01 PO4 ? N/A P40485 22574179
    1619 4a6h:D (1.449) BS02 PO4 ? N/A P40485 22574179
    1620 4a6h:D (1.449) BS03 PO4 ? N/A P40485 22574179
    1621 4a6k:A (1.8) BS01 PO4 ? N/A P40485 22574179
    1622 4a6k:B (1.8) BS01 PO4 ? N/A P40485 22574179
    1623 4a6k:C (1.8) BS01 PO4 ? N/A P40485 22574179
    1624 4a6k:C (1.8) BS02 PO4 ? N/A P40485 22574179
    1625 4a6k:D (1.8) BS01 PO4 ? N/A P40485 22574179
    1626 4a6k:D (1.8) BS02 PO4 ? N/A P40485 22574179
    1627 4a6k:D (1.8) BS03 PO4 ? N/A P40485 22574179
    1628 4ads:A (3.61) BS01 PO4 N/A GO:0042819 ... N/A 22244765
    1629 4ads:B (3.61) BS01 PO4 N/A GO:0042819 ... N/A 22244765
    1630 4ads:C (3.61) BS01 PO4 N/A GO:0042819 ... N/A 22244765
    1631 4ads:D (3.61) BS01 PO4 N/A GO:0042819 ... N/A 22244765
    1632 4ads:E (3.61) BS01 PO4 N/A GO:0042819 ... N/A 22244765
    1633 4ads:F (3.61) BS01 PO4 N/A GO:0042819 ... N/A 22244765
    1634 4adt:A (2.42) BS01 PO4 N/A GO:0042819 ... N/A 22244765
    1635 4adt:B (2.42) BS01 PO4 N/A GO:0042819 ... N/A 22244765
    1636 4ag0:A (2.304) BS01 PO4 3.1.4.52 N/A Q9HUK6 23285035
    1637 4ag0:A (2.304) BS02 PO4 3.1.4.52 N/A Q9HUK6 23285035
    1638 4ag0:B (2.304) BS01 PO4 3.1.4.52 N/A Q9HUK6 23285035
    1639 4alf:A (1.25) BS01 PO4 3.1.3.1 GO:0046872 ... Q3K5N8 25190793
    1640 4alf:B (1.25) BS01 PO4 3.1.3.1 GO:0046872 ... Q3K5N8 25190793
    1641 4aop:A (1.8) BS01 PO4 1.8.1.2 GO:0004783 ... P17846 9315848
    1642 4arv:A (1.67) BS01 PO4 3.1.3.26 N/A H9TUK6 23741456
    1643 4arv:A (1.67) BS02 PO4 3.1.3.26 N/A H9TUK6 23741456
    1644 4arv:B (1.67) BS01 PO4 3.1.3.26 N/A H9TUK6 23741456
    1645 4arv:B (1.67) BS02 PO4 3.1.3.26 N/A H9TUK6 23741456
    1646 4as4:A (1.7) BS01 PO4 3.1.3.25
    3.1.3.94    		 GO:0000287 ... P29218 23027737
    1647 4as4:B (1.7) BS01 PO4 3.1.3.25
    3.1.3.94    		 GO:0000287 ... P29218 23027737
    1648 4as5:A (2.43) BS01 PO4 3.1.3.25
    3.1.3.94    		 GO:0000287 ... O55023 23027737
    1649 4as5:B (2.43) BS01 PO4 3.1.3.25
    3.1.3.94    		 GO:0000287 ... O55023 23027737
    1650 4as5:C (2.43) BS01 PO4 3.1.3.25
    3.1.3.94    		 GO:0000287 ... O55023 23027737
    1651 4as5:D (2.43) BS01 PO4 3.1.3.25
    3.1.3.94    		 GO:0000287 ... O55023 23027737
    1652 4asj:A (2.25) BS02 PO4 2.7.7.24 GO:0000166 ... Q9HU22 23138692
    1653 4asj:C (2.25) BS02 PO4 2.7.7.24 GO:0000166 ... Q9HU22 23138692
    1654 4aui:A (3.2) BS02 PO4 ? GO:0009279 ... Q51056 23095086
    1655 4aui:A (3.2) BS03 PO4 ? GO:0009279 ... Q51056 23095086
    1656 4aui:B (3.2) BS02 PO4 ? GO:0009279 ... Q51056 23095086
    1657 4aui:B (3.2) BS03 PO4 ? GO:0009279 ... Q51056 23095086
    1658 4aui:C (3.2) BS03 PO4 ? GO:0009279 ... Q51056 23095086
    1659 4aui:C (3.2) BS04 PO4 ? GO:0009279 ... Q51056 23095086
    1660 4av6:A (4.0) BS01 PO4 7.2.3.1 GO:0000287 ... Q9S5X0 22837527
    1661 4av6:A (4.0) BS02 PO4 7.2.3.1 GO:0000287 ... Q9S5X0 22837527
    1662 4av6:B (4.0) BS01 PO4 7.2.3.1 GO:0000287 ... Q9S5X0 22837527
    1663 4av6:B (4.0) BS02 PO4 7.2.3.1 GO:0000287 ... Q9S5X0 22837527
    1664 4awn:A (1.95) BS03 PO4 3.1.21.1 GO:0002283 ... P24855 23215638
    1665 4b2o:A (1.64) BS01 PO4 3.1.4.16 GO:0004113 ... O31775 24163345
    1666 4b2o:B (1.64) BS01 PO4 3.1.4.16 GO:0004113 ... O31775 24163345
    1667 4b2o:C (1.64) BS01 PO4 3.1.4.16 GO:0004113 ... O31775 24163345
    1668 4b2o:D (1.64) BS01 PO4 3.1.4.16 GO:0004113 ... O31775 24163345
    1669 4b56:A (3.0) BS04 PO4 3.1.4.1
    3.6.1.9    		 GO:0000902 ... P06802 23041369
    1670 4b56:B (3.0) BS04 PO4 3.1.4.1
    3.6.1.9    		 GO:0000902 ... P06802 23041369
    1671 4ba1:B (1.8) BS01 PO4 3.1.13.- GO:0000175 ... Q9UXC2 23319881
    1672 4ba2:B (2.501) BS02 PO4 3.1.13.- GO:0000175 ... Q9UXC2 23319881
    1673 4bba:A (1.92) BS01 PO4 ? GO:0004857 ... Q14397 23621087
    1674 4ber:A (2.6) BS02 PO4 2.7.1.- N/A Q5ZXN6 23572077
    1675 4ber:A (2.6) BS03 PO4 2.7.1.- N/A Q5ZXN6 23572077
    1676 4ber:B (2.6) BS02 PO4 2.7.1.- N/A Q5ZXN6 23572077
    1677 4bhd:B (2.83) BS01 PO4 2.1.2.-
    4.1.2.49    		 GO:0004372 ... Q58992 25257552
    1678 4brq:A (1.45) BS02 PO4 3.6.1.5 GO:0000166 ... Q5ZUA2 23830739
    1679 4brq:A (1.45) BS03 PO4 3.6.1.5 GO:0000166 ... Q5ZUA2 23830739
    1680 4brq:B (1.45) BS02 PO4 3.6.1.5 GO:0000166 ... Q5ZUA2 23830739
    1681 4brq:B (1.45) BS03 PO4 3.6.1.5 GO:0000166 ... Q5ZUA2 23830739
    1682 4bvq:A (1.9) BS01 PO4 3.5.2.15 GO:0016787 ... P58329 23651355
    1683 4bvq:B (1.9) BS01 PO4 3.5.2.15 GO:0016787 ... P58329 23651355
    1684 4chi:A (1.27) BS02 PO4 2.6.1.- GO:0003824 ... Q4WH08 24699652
    1685 4chi:B (1.27) BS02 PO4 2.6.1.- GO:0003824 ... Q4WH08 24699652
    1686 4cmn:A (3.13) BS01 PO4 3.1.3.36
    3.1.3.56
    3.1.3.86    		 GO:0016791 ... Q01968 24704254
    1687 4cny:A (1.2) BS01 PO4 ? GO:0005524 ... P70731 24846646
    1688 4cny:A (1.2) BS02 PO4 ? GO:0005524 ... P70731 24846646
    1689 4cny:A (1.2) BS03 PO4 ? GO:0005524 ... P70731 24846646
    1690 4co4:A (1.5) BS02 PO4 ? GO:0005524 ... P70731 24846646
    1691 4co4:A (1.5) BS03 PO4 ? GO:0005524 ... P70731 24846646
    1692 4co4:B (1.5) BS01 PO4 ? GO:0005524 ... P70731 24846646
    1693 4co4:B (1.5) BS02 PO4 ? GO:0005524 ... P70731 24846646
    1694 4co4:C (1.5) BS02 PO4 ? GO:0005524 ... P70731 24846646
    1695 4co4:C (1.5) BS03 PO4 ? GO:0005524 ... P70731 24846646
    1696 4cs4:A (1.349) BS04 PO4 6.1.1.26 GO:0000049 ... Q8PWY1 24737732
    1697 4cxv:A (2.0) BS01 PO4 3.1.30.1 GO:0000014 ... Q9C9G4 25157844
    1698 4cxv:B (2.0) BS01 PO4 3.1.30.1 GO:0000014 ... Q9C9G4 25157844
    1699 4d8u:A (3.3) BS01 PO4 4.4.1.15 GO:0003824 ... Q8ZNT7 22574144
    1700 4d8u:B (3.3) BS01 PO4 4.4.1.15 GO:0003824 ... Q8ZNT7 22574144
    1701 4d8u:C (3.3) BS01 PO4 4.4.1.15 GO:0003824 ... Q8ZNT7 22574144
    1702 4d8u:D (3.3) BS01 PO4 4.4.1.15 GO:0003824 ... Q8ZNT7 22574144
    1703 4d8u:E (3.3) BS01 PO4 4.4.1.15 GO:0003824 ... Q8ZNT7 22574144
    1704 4d8u:F (3.3) BS01 PO4 4.4.1.15 GO:0003824 ... Q8ZNT7 22574144
    1705 4d8u:G (3.3) BS01 PO4 4.4.1.15 GO:0003824 ... Q8ZNT7 22574144
    1706 4d8u:H (3.3) BS01 PO4 4.4.1.15 GO:0003824 ... Q8ZNT7 22574144
    1707 4d9k:A (2.19) BS01 PO4 4.3.1.15 GO:0008838 ... P66899 22505717
    1708 4d9k:B (2.19) BS01 PO4 4.3.1.15 GO:0008838 ... P66899 22505717
    1709 4d9k:C (2.19) BS01 PO4 4.3.1.15 GO:0008838 ... P66899 22505717
    1710 4da7:A (2.05) BS02 PO4 2.4.2.1 GO:0003824 ... O34925 22957058
    1711 4dec:A (1.98) BS03 PO4 2.4.1.266 GO:0000287 ... P9WMW9 22637481
    1712 4df6:A (2.29) BS01 PO4 3.5.2.6 GO:0005576 ... P9WKD3 22587688
    1713 4dh5:A (2.2) BS04 PO4 2.7.11.11 GO:0000122 ... P05132 22751671
    1714 4dl8:A (1.698) BS03 PO4 3.6.1.23 GO:0004170 ... Q57ZH3 24001052
    1715 4dlk:A (2.02) BS05 PO4 6.3.4.18 GO:0004638 ... A0A6L8P0X7 25372694
    1716 4dpk:A (2.05) BS01 PO4 1.2.1.75 GO:0003723 ... Q96YK1 23325803
    1717 4dpk:A (2.05) BS02 PO4 1.2.1.75 GO:0003723 ... Q96YK1 23325803
    1718 4dpk:A (2.05) BS03 PO4 1.2.1.75 GO:0003723 ... Q96YK1 23325803
    1719 4dpk:B (2.05) BS01 PO4 1.2.1.75 GO:0003723 ... Q96YK1 23325803
    1720 4dpk:B (2.05) BS02 PO4 1.2.1.75 GO:0003723 ... Q96YK1 23325803
    1721 4dpk:B (2.05) BS03 PO4 1.2.1.75 GO:0003723 ... Q96YK1 23325803
    1722 4dpk:C (2.05) BS01 PO4 1.2.1.75 GO:0003723 ... Q96YK1 23325803
    1723 4dpk:C (2.05) BS02 PO4 1.2.1.75 GO:0003723 ... Q96YK1 23325803
    1724 4dpk:D (2.05) BS01 PO4 1.2.1.75 GO:0003723 ... Q96YK1 23325803
    1725 4dpk:D (2.05) BS02 PO4 1.2.1.75 GO:0003723 ... Q96YK1 23325803
    1726 4dpk:D (2.05) BS03 PO4 1.2.1.75 GO:0003723 ... Q96YK1 23325803
    1727 4dwq:A (2.25) BS03 PO4 3.1.-.-
    6.5.1.8    		 GO:0006396 ... O59245 22949672
    1728 4dwq:B (2.25) BS03 PO4 3.1.-.-
    6.5.1.8    		 GO:0006396 ... O59245 22949672
    1729 4dwq:B (2.25) BS04 PO4 3.1.-.-
    6.5.1.8    		 GO:0006396 ... O59245 22949672
    1730 4e84:A (2.6) BS03 PO4 ? GO:0003824 ... B4EB35 23256532
    1731 4e84:B (2.6) BS03 PO4 ? GO:0003824 ... B4EB35 23256532
    1732 4e8y:B (2.6) BS02 PO4 ? GO:0003824 ... B4EB35 23256532
    1733 4ear:A (1.7) BS02 PO4 2.4.2.1 GO:0000255 ... P00491 23438750
    1734 4ear:B (1.7) BS02 PO4 2.4.2.1 GO:0000255 ... P00491 23438750
    1735 4ear:C (1.7) BS02 PO4 2.4.2.1 GO:0000255 ... P00491 23438750
    1736 4eb8:A (2.3) BS01 PO4 2.4.2.1 GO:0000255 ... P00491 23438750
    1737 4eb8:B (2.3) BS01 PO4 2.4.2.1 GO:0000255 ... P00491 23438750
    1738 4eb8:C (2.3) BS01 PO4 2.4.2.1 GO:0000255 ... P00491 23438750
    1739 4eno:A (2.8) BS01 PO4 2.7.4.6 GO:0000287 ... P15531 23519676
    1740 4eno:B (2.8) BS01 PO4 2.7.4.6 GO:0000287 ... P15531 23519676
    1741 4ex7:A (1.5) BS02 PO4 ? GO:0003824 ... B6SEG4 23297194
    1742 4f0h:A (1.96) BS01 PO4 4.1.1.39 GO:0000287 ... P23755 23112176
    1743 4f0h:A (1.96) BS02 PO4 4.1.1.39 GO:0000287 ... P23755 23112176
    1744 4f5a:A (1.8) BS01 PO4 2.7.10.2 N/A P12931 23012655
    1745 4fak:A (1.7) BS02 PO4 2.1.1.177 GO:0005737 ... P0C1V0 23150671
    1746 4fdt:A (1.93) BS01 PO4 ? GO:0016787 ... Q89YI8 24529702
    1747 4fdt:B (1.93) BS01 PO4 ? GO:0016787 ... Q89YI8 24529702
    1748 4ff7:B (1.86) BS01 PO4 5.3.1.1 GO:0004807 ... P00942 22949845
    1749 4ffl:A (1.5) BS03 PO4 6.3.2.59 GO:0005524 ... Q46E79 22985965
    1750 4ffp:A (2.0) BS03 PO4 6.3.2.59 GO:0005524 ... Q46E79 22985965
    1751 4fxj:A (2.9) BS01 PO4 2.7.10.2
    2.7.11.1
    2.7.1.40    		 GO:0000287 ... P14618 23530218
    1752 4fxj:A (2.9) BS02 PO4 2.7.10.2
    2.7.11.1
    2.7.1.40    		 GO:0000287 ... P14618 23530218
    1753 4fxj:B (2.9) BS01 PO4 2.7.10.2
    2.7.11.1
    2.7.1.40    		 GO:0000287 ... P14618 23530218
    1754 4fxj:C (2.9) BS01 PO4 2.7.10.2
    2.7.11.1
    2.7.1.40    		 GO:0000287 ... P14618 23530218
    1755 4fxj:D (2.9) BS01 PO4 2.7.10.2
    2.7.11.1
    2.7.1.40    		 GO:0000287 ... P14618 23530218
    1756 4fye:A (2.413) BS01 PO4 3.1.3.67 N/A Q5ZSD5 22872863
    1757 4fyf:A (2.424) BS01 PO4 3.1.3.67 N/A Q5ZSD5 22872863
    1758 4fyv:A (2.0976) BS01 PO4 2.1.3.2 GO:0004070 ... P0A786 22906065
    1759 4g61:A (2.3) BS03 PO4 N/A N/A N/A 25263816
    1760 4g61:B (2.3) BS01 PO4 N/A N/A N/A 25263816
    1761 4ga3:A (2.39) BS04 PO4 2.5.1.1
    2.5.1.10    		 GO:0004659 ... P14324 23610597
    1762 4ga3:A (2.39) BS05 PO4 2.5.1.1
    2.5.1.10    		 GO:0004659 ... P14324 23610597
    1763 4ga4:A (3.51) BS01 PO4 2.4.2.57 GO:0004645 ... Q5JCX3 23659790
    1764 4ga4:B (3.51) BS01 PO4 2.4.2.57 GO:0004645 ... Q5JCX3 23659790
    1765 4gbd:A (1.975) BS04 PO4 ? GO:0004000 ... Q9HZ64 23050701
    1766 4gbd:B (1.975) BS04 PO4 ? GO:0004000 ... Q9HZ64 23050701
    1767 4gka:A (2.2) BS01 PO4 2.4.2.1 GO:0000255 ... P00491 23438750
    1768 4gka:B (2.2) BS01 PO4 2.4.2.1 GO:0000255 ... P00491 23438750
    1769 4gka:C (2.2) BS01 PO4 2.4.2.1 GO:0000255 ... P00491 23438750
    1770 4gka:D (2.2) BS01 PO4 2.4.2.1 GO:0000255 ... P00491 23438750
    1771 4gka:E (2.2) BS01 PO4 2.4.2.1 GO:0000255 ... P00491 23438750
    1772 4gka:F (2.2) BS01 PO4 2.4.2.1 GO:0000255 ... P00491 23438750
    1773 4gmk:A (1.72) BS01 PO4 5.3.1.6 GO:0004751 ... Q1WR87 23192019
    1774 4gmk:A (1.72) BS02 PO4 5.3.1.6 GO:0004751 ... Q1WR87 23192019
    1775 4gmk:B (1.72) BS01 PO4 5.3.1.6 GO:0004751 ... Q1WR87 23192019
    1776 4grz:A (1.3732) BS01 PO4 3.1.3.48 GO:0004725 ... P29350 23296072
    1777 4h1s:A (2.2) BS03 PO4 3.1.3.35
    3.1.3.5
    3.1.3.89
    3.1.3.91
    3.1.3.99    		 GO:0000166 ... P21589 22997138
    1778 4h1s:B (2.2) BS03 PO4 3.1.3.35
    3.1.3.5
    3.1.3.89
    3.1.3.91
    3.1.3.99    		 GO:0000166 ... P21589 22997138
    1779 4h5c:F (2.02) BS02 PO4 2.5.1.1
    2.5.1.10    		 GO:0004659 ... P14324 23234314
    1780 4h5n:A (1.86) BS01 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1781 4h5n:A (1.86) BS02 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1782 4h5n:A (1.86) BS03 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1783 4h5n:A (1.86) BS04 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1784 4h5n:A (1.86) BS05 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1785 4h5n:B (1.86) BS01 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1786 4h5n:B (1.86) BS02 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1787 4h5n:B (1.86) BS03 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1788 4h5n:B (1.86) BS04 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1789 4h5n:B (1.86) BS05 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1790 4h5n:B (1.86) BS06 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1791 4h5r:A (1.64) BS01 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1792 4h5r:A (1.64) BS02 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1793 4h5r:A (1.64) BS03 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1794 4h5r:A (1.64) BS04 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1795 4h5r:B (1.64) BS01 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1796 4h5r:B (1.64) BS02 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1797 4h5t:A (1.9) BS02 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1798 4h5w:A (1.94) BS01 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1799 4h5w:A (1.94) BS02 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    1800 4h5w:B (1.94) BS01 PO4 3.6.4.10 GO:0005524 ... P11142 25433210
    Reference:
  • Chengxin Zhang, Xi Zhang, Peter L Freddolino, and Yang Zhang. BioLiP2: an updated structure database for biologically relevent ligand-protein interactions, Nucleic Acids Research, gkad630 (2023).
  • Jianyi Yang, Ambrish Roy, and Yang Zhang. BioLiP: a semi-manually curated database for biologically relevant ligand-protein interactions, Nucleic Acids Research, 41: D1096-D1103 (2013) (download the PDF file).

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