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BioLiP
Download all results in tab-seperated text for 9664 receptor-ligand interactions, whose format is explained at readme.txt.
  • Hover over PDB to view the title of the structure. Click PDB to view the structure at the RCSB PDB database. Resolution -1.00 means the resolution is unavailable, e.g., for NMR structures.
  • Click Site # to view the binding site structure. Hover over Site # to view the binding residues.
  • Hover over Ligand to view the full ligand name. Click Ligand to view the 2D diagram and other detail information of the ligand.
  • Hover over EC number to view the full name of enzymatic activity.
  • Hover over GO terms to view all GO terms.
  • Hover over UniProt to view the protein name.
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    # PDB
    (Resolution Å)
    Site # Ligand EC number GO terms UniProt PubMed Binding
    affinity
    2801 5tau:I (6.2) BS01 ATP ? GO:0005216 ... P11716 27662087
    2802 5tav:B (4.8) BS01 ATP ? GO:0005216 ... P11716 27662087
    2803 5tav:E (4.8) BS01 ATP ? GO:0005216 ... P11716 27662087
    2804 5tav:G (4.8) BS01 ATP ? GO:0005216 ... P11716 27662087
    2805 5tav:I (4.8) BS01 ATP ? GO:0005216 ... P11716 27662087
    2806 5tc3:A (2.462) BS02 ATP 1.1.1.205 GO:0000166 ... Q756Z6 28572600
    2807 5tc3:B (2.462) BS02 ATP 1.1.1.205 GO:0000166 ... Q756Z6 28572600
    2808 5tf7:A (1.931) BS02 ATP 5.6.1.6 GO:0005524 ... P13569 N/A
    2809 5tgc:A (3.245) BS01 ATP ? GO:0000785 ... Q12406 29809203 PDBbind: -logKd/Ki=6.70, Kd=200nM
    2810 5tgc:D (3.245) BS01 ATP ? GO:0000785 ... Q12406 29809203
    2811 5tky:A (2.6) BS01 ATP 3.6.4.10 GO:0005524 ... G0SCU5 27882919
    2812 5tky:B (2.6) BS01 ATP 3.6.4.10 GO:0005524 ... G0SCU5 27882919
    2813 5tt6:A (2.187) BS01 ATP 6.5.1.3 GO:0003972 ... P00971 28223499
    2814 5tvu:A (3.5) BS01 ATP ? GO:0000287 ... F1Q9X9 28742020
    2815 5tvu:B (3.5) BS01 ATP ? GO:0000287 ... F1Q9X9 28742020
    2816 5tvw:B (2.5) BS01 ATP ? GO:0000287 ... F1Q9X9 28742020
    2817 5twv:A (6.3) BS01 ATP ? GO:0001508 ... P70673 28092267
    2818 5twv:A (6.3) BS02 ATP ? GO:0001508 ... P70673 28092267
    2819 5twv:C (6.3) BS01 ATP ? GO:0001508 ... P70673 28092267
    2820 5twv:C (6.3) BS02 ATP ? GO:0001508 ... P70673 28092267
    2821 5twv:E (6.3) BS01 ATP ? GO:0001508 ... P70673 28092267
    2822 5twv:E (6.3) BS02 ATP ? GO:0001508 ... P70673 28092267
    2823 5twv:G (6.3) BS01 ATP ? GO:0001508 ... P70673 28092267
    2824 5twv:G (6.3) BS02 ATP ? GO:0001508 ... P70673 28092267
    2825 5u03:A (6.1) BS01 ATP 6.3.4.2 GO:0002376 ... P17812 28459447
    2826 5u03:B (6.1) BS02 ATP 6.3.4.2 GO:0002376 ... P17812 28459447
    2827 5u03:C (6.1) BS01 ATP 6.3.4.2 GO:0002376 ... P17812 28459447
    2828 5u03:D (6.1) BS02 ATP 6.3.4.2 GO:0002376 ... P17812 28459447
    2829 5u8s:2 (6.1) BS02 ATP 3.6.4.12 GO:0000727 ... P29469 28096349
    2830 5u8s:2 (6.1) BS03 ATP 3.6.4.12 GO:0000727 ... P29469 28096349
    2831 5u8s:3 (6.1) BS02 ATP 3.6.4.12 GO:0000727 ... P24279 28096349
    2832 5u8s:5 (6.1) BS02 ATP 3.6.4.12 GO:0000727 ... P29496 28096349
    2833 5u8s:5 (6.1) BS03 ATP 3.6.4.12 GO:0000727 ... P29496 28096349
    2834 5u8s:6 (6.1) BS02 ATP 3.6.4.12 GO:0000727 ... P53091 28096349
    2835 5u99:A (2.4) BS01 ATP 2.4.2.17 GO:0000105 ... P9WMN1 28872824
    2836 5ub7:A (2.48) BS01 ATP N/A N/A N/A 29728456
    2837 5ub7:B (2.48) BS01 ATP N/A N/A N/A 29728456
    2838 5ubh:A (2.0) BS03 ATP 2.4.2.17 GO:0000105 ... Q5HSJ4 29208762
    2839 5ubh:B (2.0) BS02 ATP 2.4.2.17 GO:0000105 ... Q5HSJ4 29208762
    2840 5ubo:A (2.39) BS01 ATP 3.6.4.- GO:0000287 ... P68135 29259197
    2841 5uds:A (2.37) BS01 ATP 4.4.1.37 GO:0005524 ... F9UST4 28784764
    2842 5uds:B (2.37) BS01 ATP 4.4.1.37 GO:0005524 ... F9UST4 28784764
    2843 5uds:C (2.37) BS01 ATP 4.4.1.37 GO:0005524 ... F9UST4 28784764
    2844 5uds:D (2.37) BS01 ATP 4.4.1.37 GO:0005524 ... F9UST4 28784764
    2845 5uds:E (2.37) BS01 ATP 4.4.1.37 GO:0005524 ... F9UST4 28784764
    2846 5uds:F (2.37) BS01 ATP 4.4.1.37 GO:0005524 ... F9UST4 28784764
    2847 5uj7:A (3.394) BS01 ATP ? GO:0005524 ... Q13415 28112645
    2848 5uj7:B (3.394) BS01 ATP ? GO:0005524 ... Q13415 28112645
    2849 5uj7:C (3.394) BS01 ATP ? GO:0000166 ... O43929 28112645
    2850 5uj7:D (3.394) BS01 ATP ? GO:0000166 ... O43929 28112645
    2851 5uj7:E (3.394) BS01 ATP ? GO:0000808 ... O43913 28112645
    2852 5uj7:F (3.394) BS01 ATP ? GO:0000808 ... O43913 28112645
    2853 5ujm:A (18.0) BS01 ATP ? GO:0005524 ... Q13415 28112645
    2854 5ujm:D (18.0) BS01 ATP ? GO:0000166 ... O43929 28112645
    2855 5ujm:E (18.0) BS01 ATP ? GO:0000166 ... O43913 28112645
    2856 5v2l:A (2.1) BS01 ATP 4.1.1.33 GO:0004163 ... Q837E0 29025876 PDBbind: -logKd/Ki=3.54, Kd=288uM
    2857 5vc7:A (3.9) BS01 ATP ? GO:0005524 ... O05209 28390173
    2858 5vc7:A (3.9) BS02 ATP ? GO:0005524 ... O05209 28390173
    2859 5vc7:C (3.9) BS01 ATP ? GO:0005524 ... O05209 28390173
    2860 5vc7:C (3.9) BS02 ATP ? GO:0005524 ... O05209 28390173
    2861 5vc7:D (3.9) BS01 ATP ? GO:0005524 ... O05209 28390173
    2862 5vc7:D (3.9) BS02 ATP ? GO:0005524 ... O05209 28390173
    2863 5vc7:E (3.9) BS01 ATP ? GO:0005524 ... O05209 28390173
    2864 5vc7:E (3.9) BS02 ATP ? GO:0005524 ... O05209 28390173
    2865 5vc7:F (3.9) BS01 ATP ? GO:0005524 ... O05209 28390173
    2866 5vc7:F (3.9) BS02 ATP ? GO:0005524 ... O05209 28390173
    2867 5vc7:G (3.9) BS01 ATP ? GO:0005524 ... O05209 28390173
    2868 5vc7:G (3.9) BS02 ATP ? GO:0005524 ... O05209 28390173
    2869 5vqa:A (2.54) BS01 ATP ? GO:0001556 ... Q15645 28659378
    2870 5w81:A (3.37) BS01 ATP 5.6.1.6 GO:0002679 ... Q1LX78 28735752
    2871 5w81:A (3.37) BS02 ATP 5.6.1.6 GO:0002679 ... Q1LX78 28735752
    2872 5wc0:A (4.4) BS01 ATP 5.6.1.1 GO:0000212 ... P34808 28783150
    2873 5wc0:B (4.4) BS01 ATP 5.6.1.1 GO:0000212 ... P34808 28783150
    2874 5wc0:B (4.4) BS02 ATP 5.6.1.1 GO:0000212 ... P34808 28783150
    2875 5wc0:C (4.4) BS01 ATP 5.6.1.1 GO:0000212 ... P34808 28783150
    2876 5wc0:C (4.4) BS02 ATP 5.6.1.1 GO:0000212 ... P34808 28783150
    2877 5wc0:D (4.4) BS01 ATP 5.6.1.1 GO:0000212 ... P34808 28783150
    2878 5wc0:D (4.4) BS02 ATP 5.6.1.1 GO:0000212 ... P34808 28783150
    2879 5wc0:E (4.4) BS01 ATP 5.6.1.1 GO:0000212 ... P34808 28783150
    2880 5wc0:E (4.4) BS02 ATP 5.6.1.1 GO:0000212 ... P34808 28783150
    2881 5wc0:F (4.4) BS01 ATP 5.6.1.1 GO:0000212 ... P34808 28783150
    2882 5wc0:F (4.4) BS02 ATP 5.6.1.1 GO:0000212 ... P34808 28783150
    2883 5wcb:B (6.0) BS01 ATP 5.6.1.1 GO:0000212 ... P34808 28783150
    2884 5wcb:C (6.0) BS01 ATP 5.6.1.1 GO:0000212 ... P34808 28783150
    2885 5wcb:D (6.0) BS01 ATP 5.6.1.1 GO:0000212 ... P34808 28783150
    2886 5wcb:E (6.0) BS01 ATP 5.6.1.1 GO:0000212 ... P34808 28783150
    2887 5wcb:F (6.0) BS01 ATP 5.6.1.1 GO:0000212 ... P34808 28783150
    2888 5wni:A (2.65) BS02 ATP 2.7.11.1 GO:0004672 ... Q9ERK0 29706531
    2889 5wrs:A (2.75) BS01 ATP ? N/A Q96MK3 28432788 PDBbind: -logKd/Ki=5.52, Kd=3uM
    2890 5wrs:B (2.75) BS01 ATP ? N/A Q96MK3 28432788 PDBbind: -logKd/Ki=5.52, Kd=3uM
    2891 5ws9:A (1.9) BS04 ATP 2.7.1.40 GO:0000287 ... P9WKE5 29215013
    2892 5ws9:B (1.9) BS04 ATP 2.7.1.40 GO:0000287 ... P9WKE5 29215013
    2893 5ws9:C (1.9) BS04 ATP 2.7.1.40 GO:0000287 ... P9WKE5 29215013
    2894 5ws9:D (1.9) BS04 ATP 2.7.1.40 GO:0000287 ... P9WKE5 29215013
    2895 5wu4:A (2.8) BS01 ATP 2.7.7.19
    2.7.7.52
    N/A Q9H6E5 28589955
    2896 5wu4:B (2.8) BS01 ATP 2.7.7.19
    2.7.7.52
    N/A Q9H6E5 28589955
    2897 5x1q:A (1.602) BS01 ATP ? GO:0004672 ... A0A1S4NYE5 29858276
    2898 5x6c:A (3.101) BS03 ATP 6.1.1.27 GO:0000049 ... Q59054 29142195
    2899 5x6c:B (3.101) BS03 ATP 6.1.1.27 GO:0000049 ... Q59054 29142195
    2900 5x8a:B (2.51) BS01 ATP 2.7.4.9 GO:0004798 ... Q5SHX3 29652261
    2901 5x8b:A (1.39) BS01 ATP 2.7.4.9 GO:0004798 ... Q5SHX3 29652261
    2902 5x9g:A (3.0) BS01 ATP ? GO:0015095 ... Q5SMG8 28747715
    2903 5x9g:B (3.0) BS01 ATP ? GO:0015095 ... Q5SMG8 28747715
    2904 5x9g:C (3.0) BS01 ATP ? GO:0015095 ... Q5SMG8 28747715
    2905 5x9g:D (3.0) BS01 ATP ? GO:0015095 ... Q5SMG8 28747715
    2906 5x9h:A (3.598) BS01 ATP ? GO:0000287 ... Q5SMG8 28747715 PDBbind: -logKd/Ki=3.76, Kd=172uM
    2907 5x9h:B (3.598) BS01 ATP ? GO:0000287 ... Q5SMG8 28747715 PDBbind: -logKd/Ki=3.76, Kd=172uM
    2908 5xd1:A (1.6) BS02 ATP 3.6.1.58
    3.6.1.61
    3.6.1.69
    GO:0006260 ... A0QUZ2 28705712
    2909 5xd2:A (1.75) BS02 ATP 3.6.1.58
    3.6.1.61
    3.6.1.69
    GO:0006260 ... A0QUZ2 28705712
    2910 5xd3:A (1.78) BS01 ATP 3.6.1.58
    3.6.1.61
    3.6.1.69
    GO:0006260 ... A0QUZ2 28705712
    2911 5xd3:A (1.78) BS02 ATP 3.6.1.58
    3.6.1.61
    3.6.1.69
    GO:0006260 ... A0QUZ2 28705712
    2912 5xd4:A (1.47) BS01 ATP 3.6.1.58
    3.6.1.61
    3.6.1.69
    GO:0006260 ... A0QUZ2 28705712
    2913 5xd4:A (1.47) BS02 ATP 3.6.1.58
    3.6.1.61
    3.6.1.69
    GO:0006260 ... A0QUZ2 28705712
    2914 5xd5:A (1.75) BS01 ATP 3.6.1.58
    3.6.1.61
    3.6.1.69
    GO:0006260 ... A0QUZ2 28705712
    2915 5xd5:B (1.75) BS01 ATP 3.6.1.58
    3.6.1.61
    3.6.1.69
    GO:0006260 ... A0QUZ2 28705712
    2916 5xjc:Q (3.6) BS02 ATP 3.6.4.13 GO:0000398 ... O60306 28502770
    2917 5xjc:u (3.6) BS03 ATP 3.6.4.13 GO:0000184 ... P38919 28502770
    2918 5xmi:B (3.9) BS01 ATP ? GO:0001778 ... P52917 28714467
    2919 5xmi:C (3.9) BS01 ATP ? GO:0001778 ... P52917 28714467
    2920 5xmi:C (3.9) BS02 ATP ? GO:0001778 ... P52917 28714467
    2921 5xmi:D (3.9) BS01 ATP ? GO:0001778 ... P52917 28714467
    2922 5xmi:D (3.9) BS02 ATP ? GO:0001778 ... P52917 28714467
    2923 5xmi:E (3.9) BS01 ATP ? GO:0001778 ... P52917 28714467
    2924 5xmi:E (3.9) BS02 ATP ? GO:0001778 ... P52917 28714467
    2925 5xmi:F (3.9) BS01 ATP ? GO:0001778 ... P52917 28714467
    2926 5xmk:B (4.18) BS01 ATP ? GO:0001778 ... P52917 28714467
    2927 5xmk:C (4.18) BS01 ATP ? GO:0001778 ... P52917 28714467
    2928 5xmk:C (4.18) BS02 ATP ? GO:0001778 ... P52917 28714467
    2929 5xmk:D (4.18) BS01 ATP ? GO:0001778 ... P52917 28714467
    2930 5xmk:D (4.18) BS02 ATP ? GO:0001778 ... P52917 28714467
    2931 5xmk:E (4.18) BS01 ATP ? GO:0001778 ... P52917 28714467
    2932 5xmk:E (4.18) BS02 ATP ? GO:0001778 ... P52917 28714467
    2933 5xmk:F (4.18) BS01 ATP ? GO:0001778 ... P52917 28714467
    2934 5xvu:A (3.0) BS01 ATP 2.7.11.1 GO:0004672 ... Q8IIR9 29743645 PDBbind: -logKd/Ki=4.65, Kd=22.3uM
    2935 5xvu:B (3.0) BS01 ATP 2.7.11.1 GO:0004672 ... Q8IIR9 29743645
    2936 5xvu:C (3.0) BS01 ATP 2.7.11.1 GO:0004672 ... Q8IIR9 29743645
    2937 5xz9:A (2.0) BS01 ATP 2.7.1.11 GO:0003872 ... P99165 29940104
    2938 5y0n:A (2.302) BS01 ATP 6.3.4.- GO:0000049 ... O34513 30150682
    2939 5y0n:B (2.302) BS01 ATP 6.3.4.- GO:0000049 ... O34513 30150682
    2940 5ya1:A (2.701) BS01 ATP 2.7.1.189 GO:0005515 ... P77432 29868643
    2941 5ya1:B (2.701) BS01 ATP 2.7.1.189 GO:0005515 ... P77432 29868643
    2942 5yec:A (2.147) BS02 ATP ? GO:0008641 ... P38862 29237558
    2943 5yec:C (2.147) BS02 ATP ? GO:0008641 ... P38862 29237558
    2944 5yee:B (1.81) BS02 ATP 3.6.4.- GO:0000287 ... P68135 30283132
    2945 5yh2:A (3.55) BS01 ATP ? GO:0001934 ... Q96MK3 29572475
    2946 5yh2:B (3.55) BS01 ATP ? GO:0001934 ... Q96MK3 29572475
    2947 5yh3:A (3.3) BS01 ATP ? GO:0001934 ... Q96MK3 29572475
    2948 5yh3:A (3.3) BS02 ATP ? GO:0001934 ... Q96MK3 29572475
    2949 5yh3:B (3.3) BS01 ATP ? GO:0001934 ... Q96MK3 29572475
    2950 5yh3:B (3.3) BS02 ATP ? GO:0001934 ... Q96MK3 29572475
    2951 5ypu:A (2.0) BS02 ATP 3.6.4.- GO:0000287 ... P68135 30254171
    2952 5ypu:C (2.0) BS02 ATP 3.6.4.- GO:0000287 ... P68135 30254171
    2953 5yud:A (4.28) BS01 ATP ? GO:0005515 ... Q9R016 29182158
    2954 5yzg:u (4.1) BS03 ATP 3.6.4.13 GO:0000184 ... P38919 29301961
    2955 5z68:A (3.0) BS01 ATP ? GO:0000731 ... Q8RDL3 29391496 PDBbind: -logKd/Ki=5.89, Kd=1.3uM
    2956 5z68:A (3.0) BS02 ATP ? GO:0000731 ... Q8RDL3 29391496 PDBbind: -logKd/Ki=5.89, Kd=1.3uM
    2957 5z68:B (3.0) BS01 ATP ? GO:0000731 ... Q8RDL3 29391496 PDBbind: -logKd/Ki=5.89, Kd=1.3uM
    2958 5z68:B (3.0) BS02 ATP ? GO:0000731 ... Q8RDL3 29391496 PDBbind: -logKd/Ki=5.89, Kd=1.3uM
    2959 5z68:C (3.0) BS01 ATP ? GO:0000731 ... Q8RDL3 29391496
    2960 5z68:D (3.0) BS01 ATP ? GO:0000731 ... Q8RDL3 29391496
    2961 5z68:D (3.0) BS02 ATP ? GO:0000731 ... Q8RDL3 29391496
    2962 5z6r:A (3.0) BS01 ATP 5.6.1.1 GO:0005524 ... Q9UBP0 N/A
    2963 5zdk:A (2.45) BS01 ATP 6.1.1.18 GO:0000166 ... Q5SKU4 30248426
    2964 5zdl:A (2.6) BS01 ATP 6.1.1.18 GO:0000166 ... Q5SKU4 30248426
    2965 5zlp:A (2.93) BS01 ATP 6.3.1.2 GO:0003824 ... P94845 30076387
    2966 5zlp:B (2.93) BS01 ATP 6.3.1.2 GO:0003824 ... P94845 30076387
    2967 5zlp:C (2.93) BS01 ATP 6.3.1.2 GO:0003824 ... P94845 30076387
    2968 5zlp:F (2.93) BS01 ATP 6.3.1.2 GO:0003824 ... P94845 30076387
    2969 5zlp:G (2.93) BS01 ATP 6.3.1.2 GO:0003824 ... P94845 30076387
    2970 5zlp:K (2.93) BS01 ATP 6.3.1.2 GO:0003824 ... P94845 30076387
    2971 5ztb:A (2.2) BS04 ATP 2.8.1.15 GO:0000049 ... Q72LF3 32265486
    2972 5ztb:B (2.2) BS04 ATP 2.8.1.15 GO:0000049 ... Q72LF3 32265486
    2973 5ztb:C (2.2) BS04 ATP 2.8.1.15 GO:0000049 ... Q72LF3 32265486
    2974 5zua:A (2.471) BS01 ATP 3.4.19.12 N/A T2B9G2 N/A
    2975 5zxd:A (2.288) BS01 ATP ? GO:0005524 ... Q8NE71 30017566
    2976 5zxd:A (2.288) BS02 ATP ? GO:0005524 ... Q8NE71 30017566
    2977 5zxd:B (2.288) BS01 ATP ? GO:0005524 ... Q8NE71 30017566
    2978 5zxd:B (2.288) BS02 ATP ? GO:0005524 ... Q8NE71 30017566
    2979 5zza:A (1.53) BS03 ATP 3.6.4.- GO:0000287 ... P68135 30283132
    2980 5zzb:B (2.3) BS02 ATP 3.6.4.- GO:0000287 ... P68135 30283132
    2981 5zzb:D (2.3) BS02 ATP 3.6.4.- GO:0000287 ... P68135 30283132
    2982 6a8a:A (1.8) BS02 ATP 2.7.1.15 GO:0004747 ... A0A3S7X0F5 31170491
    2983 6a8a:B (1.8) BS02 ATP 2.7.1.15 GO:0004747 ... A0A3S7X0F5 31170491
    2984 6aac:A (1.479) BS02 ATP 6.1.1.26 GO:0000049 ... Q8PWY1 31031143
    2985 6aad:A (1.444) BS02 ATP 6.1.1.26 GO:0000049 ... Q8PWY1 31031143
    2986 6aan:A (1.51) BS02 ATP 6.1.1.26 GO:0000049 ... Q8PWY1 31031143
    2987 6aao:A (1.403) BS02 ATP 6.1.1.26 GO:0000049 ... Q8PWY1 31031143
    2988 6aap:A (1.5) BS02 ATP 6.1.1.26 GO:0000049 ... Q8PWY1 31031143
    2989 6aaq:A (1.551) BS02 ATP 6.1.1.26 GO:0000049 ... Q8PWY1 31031143
    2990 6aaz:A (1.842) BS02 ATP 6.1.1.26 GO:0000049 ... Q8PWY1 31031143
    2991 6ab0:A (1.441) BS02 ATP 6.1.1.26 GO:0000049 ... Q8PWY1 31031143
    2992 6ab1:A (1.381) BS02 ATP 6.1.1.26 GO:0000049 ... Q8PWY1 31031143
    2993 6ab2:A (1.4) BS02 ATP 6.1.1.26 GO:0000049 ... Q8PWY1 31031143
    2994 6ab8:A (1.753) BS02 ATP 6.1.1.26 GO:0000049 ... Q8PWY1 31031143
    2995 6abk:A (1.58) BS02 ATP 6.1.1.26 GO:0000049 ... Q8PWY1 31031143
    2996 6abl:A (1.47) BS02 ATP 6.1.1.26 GO:0000049 ... Q8PWY1 31031143
    2997 6abm:A (1.368) BS02 ATP 6.1.1.26 GO:0000049 ... Q8PWY1 31031143
    2998 6aec:A (1.72) BS03 ATP 2.7.7.7 GO:0003677 ... Q9NP87 31067051 MOAD: Kd=0.71uM
    PDBbind: -logKd/Ki=6.15, Kd=0.71uM
    2999 6ah4:A (3.296) BS01 ATP ? GO:0001614 ... P56373 31232692 BindingDB: EC50=340nM
    3000 6ah4:A (3.296) BS02 ATP ? GO:0001614 ... P56373 31232692 BindingDB: EC50=340nM

    Reference:
  • Chengxin Zhang, Xi Zhang, Peter L Freddolino, and Yang Zhang. BioLiP2: an updated structure database for biologically relevent ligand-protein interactions, Nucleic Acids Research, gkad630 (2023).
  • Jianyi Yang, Ambrish Roy, and Yang Zhang. BioLiP: a semi-manually curated database for biologically relevant ligand-protein interactions, Nucleic Acids Research, 41: D1096-D1103 (2013) (download the PDF file).
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