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BioLiP
Download all results in tab-seperated text for 594392 receptor-ligand interactions, whose format is explained at readme.txt.
  • Hover over PDB to view the title of the structure. Click PDB to view the structure at the RCSB PDB database. Resolution -1.00 means the resolution is unavailable, e.g., for NMR structures.
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  • Hover over Ligand to view the full ligand name. Click Ligand to view the 2D diagram and other detail information of the ligand.
  • Hover over EC number to view the full name of enzymatic activity.
  • Hover over GO terms to view all GO terms.
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    # PDB
    (Resolution Å)
    Site # Ligand EC number GO terms UniProt PubMed Binding
    affinity
    18001 1kx5:H (1.94) BS01 dna ? GO:0000786 ... P02281 12079350
    18002 1kx5:H (1.94) BS02 dna ? GO:0000786 ... P02281 12079350
    18003 1kxg:A (2.0) BS01 MG ? GO:0005164 ... Q9Y275 11862220
    18004 1kxg:F (2.0) BS01 MG ? GO:0005164 ... Q9Y275 11862220
    18005 1kxp:A (2.1) BS01 ATP 3.6.4.- GO:0000287 ... P68135 12048248
    18006 1kxq:A (1.6) BS01 CA 3.2.1.1 GO:0003824 ... P00690 11960990
    18007 1kxq:B (1.6) BS01 CA 3.2.1.1 GO:0003824 ... P00690 11960990
    18008 1kxq:C (1.6) BS01 CA 3.2.1.1 GO:0003824 ... P00690 11960990
    18009 1kxq:D (1.6) BS01 CA 3.2.1.1 GO:0003824 ... P00690 11960990
    18010 1kxt:A (2.0) BS01 CA 3.2.1.1 GO:0003824 ... P00690 11960990
    18011 1kxt:C (2.0) BS01 CA 3.2.1.1 GO:0003824 ... P00690 11960990
    18012 1kxt:E (2.0) BS01 CA 3.2.1.1 GO:0003824 ... P00690 11960990
    18013 1ky4:A (2.8) BS01 NAD 3.13.2.1 GO:0001666 ... P10760 11927587
    18014 1ky4:A (2.8) BS02 NAD 3.13.2.1 GO:0001666 ... P10760 11927587
    18015 1ky4:B (2.8) BS01 NAD 3.13.2.1 GO:0001666 ... P10760 11927587
    18016 1ky4:B (2.8) BS02 NAD 3.13.2.1 GO:0001666 ... P10760 11927587
    18017 1ky4:C (2.8) BS01 NAD 3.13.2.1 GO:0001666 ... P10760 11927587
    18018 1ky4:C (2.8) BS02 NAD 3.13.2.1 GO:0001666 ... P10760 11927587
    18019 1ky4:D (2.8) BS01 NAD 3.13.2.1 GO:0001666 ... P10760 11927587
    18020 1ky4:D (2.8) BS02 NAD 3.13.2.1 GO:0001666 ... P10760 11927587
    18021 1ky5:A (2.8) BS01 NAI 3.13.2.1 GO:0001666 ... P10760 11927587
    18022 1ky5:A (2.8) BS02 ADY 3.13.2.1 GO:0001666 ... P10760 11927587
    18023 1ky5:A (2.8) BS03 NAI 3.13.2.1 GO:0001666 ... P10760 11927587
    18024 1ky5:B (2.8) BS01 NAI 3.13.2.1 GO:0001666 ... P10760 11927587
    18025 1ky5:B (2.8) BS02 NAI 3.13.2.1 GO:0001666 ... P10760 11927587
    18026 1ky5:B (2.8) BS03 ADY 3.13.2.1 GO:0001666 ... P10760 11927587
    18027 1ky5:C (2.8) BS01 NAI 3.13.2.1 GO:0001666 ... P10760 11927587
    18028 1ky5:C (2.8) BS02 ADY 3.13.2.1 GO:0001666 ... P10760 11927587
    18029 1ky5:C (2.8) BS03 NAI 3.13.2.1 GO:0001666 ... P10760 11927587
    18030 1ky5:D (2.8) BS01 NAI 3.13.2.1 GO:0001666 ... P10760 11927587
    18031 1ky5:D (2.8) BS02 NAI 3.13.2.1 GO:0001666 ... P10760 11927587
    18032 1ky5:D (2.8) BS03 ADY 3.13.2.1 GO:0001666 ... P10760 11927587
    18033 1ky6:A (2.0) BS01 peptide ? GO:0006886 ... P17427 12057195
    18034 1ky7:A (2.15) BS01 peptide ? GO:0006886 ... P17427 12057195
    18035 1kya:A (2.4) BS01 CU 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18036 1kya:A (2.4) BS02 CU 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18037 1kya:A (2.4) BS03 CU 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18038 1kya:A (2.4) BS04 CU 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18039 1kya:A (2.4) BS05 XYD 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18040 1kya:B (2.4) BS01 CU 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18041 1kya:B (2.4) BS02 CU 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18042 1kya:B (2.4) BS03 CU 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18043 1kya:B (2.4) BS04 CU 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18044 1kya:B (2.4) BS05 XYD 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18045 1kya:C (2.4) BS01 CU 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18046 1kya:C (2.4) BS02 CU 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18047 1kya:C (2.4) BS03 CU 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18048 1kya:C (2.4) BS04 CU 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18049 1kya:C (2.4) BS05 XYD 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18050 1kya:D (2.4) BS01 CU 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18051 1kya:D (2.4) BS02 CU 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18052 1kya:D (2.4) BS03 CU 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18053 1kya:D (2.4) BS04 CU 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18054 1kya:D (2.4) BS05 XYD 1.10.3.2 GO:0005507 ... Q96UT7 12044164
    18055 1kyd:A (2.0) BS01 peptide ? GO:0006886 ... P17427 12057195
    18056 1kyf:A (1.22) BS01 peptide ? GO:0006886 ... P17427 12057195
    18057 1kyi:A (3.1) BS01 ATP ? GO:0005515 ... P43773 12054822
    18058 1kyi:B (3.1) BS01 ATP ? GO:0005515 ... P43773 12054822
    18059 1kyi:C (3.1) BS01 ATP ? GO:0005515 ... P43773 12054822
    18060 1kyi:D (3.1) BS01 ATP ? GO:0005515 ... P43773 12054822
    18061 1kyi:E (3.1) BS01 ATP ? GO:0005515 ... P43773 12054822
    18062 1kyi:F (3.1) BS01 ATP ? GO:0005515 ... P43773 12054822
    18063 1kyi:G (3.1) BS01 LVS 3.4.25.2 GO:0004298 ... P43772 12054822
    18064 1kyi:H (3.1) BS01 LVS 3.4.25.2 GO:0004298 ... P43772 12054822
    18065 1kyi:I (3.1) BS01 LVS 3.4.25.2 GO:0004298 ... P43772 12054822
    18066 1kyi:J (3.1) BS01 LVS 3.4.25.2 GO:0004298 ... P43772 12054822
    18067 1kyi:K (3.1) BS01 LVS 3.4.25.2 GO:0004298 ... P43772 12054822
    18068 1kyi:L (3.1) BS01 LVS 3.4.25.2 GO:0004298 ... P43772 12054822
    18069 1kyi:M (3.1) BS01 LVS 3.4.25.2 GO:0004298 ... P43772 12054822
    18070 1kyi:N (3.1) BS01 LVS 3.4.25.2 GO:0004298 ... P43772 12054822
    18071 1kyi:O (3.1) BS01 LVS 3.4.25.2 GO:0004298 ... P43772 12054822
    18072 1kyi:P (3.1) BS01 LVS 3.4.25.2 GO:0004298 ... P43772 12054822
    18073 1kyi:Q (3.1) BS01 LVS 3.4.25.2 GO:0004298 ... P43772 12054822
    18074 1kyi:R (3.1) BS01 LVS 3.4.25.2 GO:0004298 ... P43772 12054822
    18075 1kyi:S (3.1) BS01 ATP ? GO:0005515 ... P43773 12054822
    18076 1kyi:T (3.1) BS01 ATP ? GO:0005515 ... P43773 12054822
    18077 1kyi:U (3.1) BS01 ATP ? GO:0005515 ... P43773 12054822
    18078 1kyi:V (3.1) BS01 ATP ? GO:0005515 ... P43773 12054822
    18079 1kyi:W (3.1) BS01 ATP ? GO:0005515 ... P43773 12054822
    18080 1kyi:X (3.1) BS01 ATP ? GO:0005515 ... P43773 12054822
    18081 1kyn:A (3.5) BS01 KTP 3.4.21.20 GO:0002003 ... P08311 11942800 PDBbind: -logKd/Ki=5.39, IC50=4.1uM
    BindingDB: IC50=4100nM
    18082 1kyn:B (3.5) BS01 KTP 3.4.21.20 GO:0002003 ... P08311 11942800 BindingDB: IC50=4100nM
    18083 1kyo:C (2.97) BS01 HEC 7.1.1.8 GO:0005739 ... P00163 11880631
    18084 1kyo:C (2.97) BS02 HEC 7.1.1.8 GO:0005739 ... P00163 11880631
    18085 1kyo:C (2.97) BS03 SMA 7.1.1.8 GO:0005739 ... P00163 11880631
    18086 1kyo:E (2.97) BS01 FES 7.1.1.8 GO:0008121 ... P08067 11880631
    18087 1kyo:E (2.97) BS02 SMA 7.1.1.8 GO:0008121 ... P08067 11880631
    18088 1kyo:N (2.97) BS01 HEC 7.1.1.8 GO:0005739 ... P00163 11880631
    18089 1kyo:N (2.97) BS02 HEC 7.1.1.8 GO:0005739 ... P00163 11880631
    18090 1kyo:N (2.97) BS03 SMA 7.1.1.8 GO:0005739 ... P00163 11880631
    18091 1kyo:P (2.97) BS01 FES 7.1.1.8 GO:0008121 ... P08067 11880631
    18092 1kyo:W (2.97) BS01 HEC ? GO:0005515 ... P00044 11880631
    18093 1kyq:A (2.2) BS01 NAD 1.3.1.76
    4.99.1.4
    GO:0000103 ... P15807 11980703
    18094 1kyq:B (2.2) BS01 NAD 1.3.1.76
    4.99.1.4
    GO:0000103 ... P15807 11980703
    18095 1kyq:C (2.2) BS01 NAD 1.3.1.76
    4.99.1.4
    GO:0000103 ... P15807 11980703
    18096 1kyr:A (1.5) BS01 CU ? GO:0003674 ... P42212 11929238
    18097 1kys:A (1.44) BS01 ZN ? GO:0003674 ... P42212 11929238
    18098 1kyt:A (1.7) BS01 CA 3.1.3.18 GO:0000287 ... Q9HLQ2 N/A
    18099 1kyt:A (1.7) BS02 CA 3.1.3.18 GO:0000287 ... Q9HLQ2 N/A
    18100 1kyt:A (1.7) BS03 CA 3.1.3.18 GO:0000287 ... Q9HLQ2 N/A
    18101 1kyt:B (1.7) BS01 CA 3.1.3.18 GO:0000287 ... Q9HLQ2 N/A
    18102 1kyt:B (1.7) BS02 CA 3.1.3.18 GO:0000287 ... Q9HLQ2 N/A
    18103 1kyu:A (1.8) BS01 peptide ? GO:0006886 ... P17427 12057195
    18104 1kyv:A (2.4) BS01 RBF 2.5.1.78 GO:0000906 ... Q9UUB1 12083520 MOAD: Kd=1.2uM
    18105 1kyv:A (2.4) BS02 RBF 2.5.1.78 GO:0000906 ... Q9UUB1 12083520 MOAD: Kd=1.2uM
    18106 1kyv:B (2.4) BS01 RBF 2.5.1.78 GO:0000906 ... Q9UUB1 12083520 MOAD: Kd=1.2uM
    18107 1kyv:B (2.4) BS02 RBF 2.5.1.78 GO:0000906 ... Q9UUB1 12083520 MOAD: Kd=1.2uM
    18108 1kyv:C (2.4) BS01 RBF 2.5.1.78 GO:0000906 ... Q9UUB1 12083520 MOAD: Kd=1.2uM
    18109 1kyv:C (2.4) BS02 RBF 2.5.1.78 GO:0000906 ... Q9UUB1 12083520 MOAD: Kd=1.2uM
    18110 1kyv:D (2.4) BS01 RBF 2.5.1.78 GO:0000906 ... Q9UUB1 12083520 MOAD: Kd=1.2uM
    18111 1kyv:D (2.4) BS02 RBF 2.5.1.78 GO:0000906 ... Q9UUB1 12083520 MOAD: Kd=1.2uM
    18112 1kyv:E (2.4) BS01 RBF 2.5.1.78 GO:0000906 ... Q9UUB1 12083520 MOAD: Kd=1.2uM
    18113 1kyv:E (2.4) BS02 RBF 2.5.1.78 GO:0000906 ... Q9UUB1 12083520 MOAD: Kd=1.2uM
    18114 1kyx:A (2.6) BS01 CRM 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18115 1kyx:A (2.6) BS02 CRM 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18116 1kyx:B (2.6) BS01 CRM 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18117 1kyx:B (2.6) BS02 CRM 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18118 1kyx:C (2.6) BS01 CRM 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18119 1kyx:C (2.6) BS02 CRM 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18120 1kyx:D (2.6) BS01 CRM 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18121 1kyx:D (2.6) BS02 CRM 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18122 1kyx:E (2.6) BS01 CRM 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18123 1kyx:E (2.6) BS02 CRM 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18124 1kyy:A (2.4) BS01 INI 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18125 1kyy:A (2.4) BS02 INI 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18126 1kyy:B (2.4) BS01 INI 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18127 1kyy:B (2.4) BS02 INI 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18128 1kyy:C (2.4) BS01 INI 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18129 1kyy:C (2.4) BS02 INI 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18130 1kyy:D (2.4) BS01 INI 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18131 1kyy:D (2.4) BS02 INI 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18132 1kyy:E (2.4) BS01 INI 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18133 1kyy:E (2.4) BS02 INI 2.5.1.78 GO:0000906 ... Q9UUB1 12083520
    18134 1kz8:A (2.0) BS01 F6P 3.1.3.11 GO:0005737 ... P00636 12190310
    18135 1kz8:A (2.0) BS02 MN 3.1.3.11 GO:0005737 ... P00636 12190310
    18136 1kz8:A (2.0) BS03 AMP 3.1.3.11 GO:0005737 ... P00636 12190310 BindingDB: IC50=1300nM
    18137 1kz8:A (2.0) BS04 PFE 3.1.3.11 GO:0005737 ... P00636 12190310 MOAD: ic50=0.83uM
    PDBbind: -logKd/Ki=6.08, IC50=0.83uM
    18138 1kz8:F (2.0) BS01 F6P 3.1.3.11 GO:0005737 ... P00636 12190310
    18139 1kz8:F (2.0) BS02 MN 3.1.3.11 GO:0005737 ... P00636 12190310
    18140 1kz8:F (2.0) BS03 AMP 3.1.3.11 GO:0005737 ... P00636 12190310 BindingDB: IC50=1300nM
    18141 1kz8:F (2.0) BS04 PFE 3.1.3.11 GO:0005737 ... P00636 12190310 MOAD: ic50=0.83uM
    PDBbind: -logKd/Ki=6.08, IC50=0.83uM
    18142 1kzh:A (2.55) BS01 SO4 2.7.1.90 GO:0003872 ... P70826 12015149
    18143 1kzh:A (2.55) BS02 SO4 2.7.1.90 GO:0003872 ... P70826 12015149
    18144 1kzh:A (2.55) BS03 SO4 2.7.1.90 GO:0003872 ... P70826 12015149
    18145 1kzh:B (2.55) BS01 SO4 2.7.1.90 GO:0003872 ... P70826 12015149
    18146 1kzh:B (2.55) BS02 SO4 2.7.1.90 GO:0003872 ... P70826 12015149
    18147 1kzh:B (2.55) BS03 SO4 2.7.1.90 GO:0003872 ... P70826 12015149
    18148 1kzh:B (2.55) BS04 SO4 2.7.1.90 GO:0003872 ... P70826 12015149
    18149 1kzh:B (2.55) BS05 SO4 2.7.1.90 GO:0003872 ... P70826 12015149
    18150 1kzi:A (1.75) BS01 CO3 2.1.1.45 GO:0000287 ... P0A884 12033935
    18151 1kzi:A (1.75) BS02 UMP 2.1.1.45 GO:0000287 ... P0A884 12033935
    18152 1kzi:A (1.75) BS03 THG 2.1.1.45 GO:0000287 ... P0A884 12033935
    18153 1kzi:A (1.75) BS04 UMP 2.1.1.45 GO:0000287 ... P0A884 12033935
    18154 1kzi:B (1.75) BS01 UMP 2.1.1.45 GO:0000287 ... P0A884 12033935
    18155 1kzi:B (1.75) BS02 UMP 2.1.1.45 GO:0000287 ... P0A884 12033935
    18156 1kzi:B (1.75) BS03 THG 2.1.1.45 GO:0000287 ... P0A884 12033935
    18157 1kzj:A (2.6) BS01 UMP 2.1.1.45 GO:0000287 ... P0A884 12033935
    18158 1kzj:A (2.6) BS02 UMP 2.1.1.45 GO:0000287 ... P0A884 12033935
    18159 1kzj:B (2.6) BS01 UMP 2.1.1.45 GO:0000287 ... P0A884 12033935
    18160 1kzj:B (2.6) BS02 UMP 2.1.1.45 GO:0000287 ... P0A884 12033935
    18161 1kzj:C (2.6) BS01 UMP 2.1.1.45 GO:0000287 ... P0A884 12033935
    18162 1kzj:D (2.6) BS01 UMP 2.1.1.45 GO:0000287 ... P0A884 12033935
    18163 1kzj:D (2.6) BS02 UMP 2.1.1.45 GO:0000287 ... P0A884 12033935
    18164 1kzj:E (2.6) BS01 UMP 2.1.1.45 GO:0000287 ... P0A884 12033935
    18165 1kzj:E (2.6) BS02 UMP 2.1.1.45 GO:0000287 ... P0A884 12033935
    18166 1kzj:F (2.6) BS01 UMP 2.1.1.45 GO:0000287 ... P0A884 12033935
    18167 1kzl:A (2.1) BS01 CRM 2.5.1.9 GO:0004746 ... Q9Y7P0 12377123
    18168 1kzl:A (2.1) BS02 CRM 2.5.1.9 GO:0004746 ... Q9Y7P0 12377123
    18169 1kzm:A (2.0) BS01 CA 1.11.1.7 GO:0004601 ... P00433 12351824
    18170 1kzm:A (2.0) BS02 CA 1.11.1.7 GO:0004601 ... P00433 12351824
    18171 1kzm:A (2.0) BS03 HEM 1.11.1.7 GO:0004601 ... P00433 12351824
    18172 1kzo:B (2.2) BS01 peptide 2.5.1.58 GO:0003824 ... Q02293 12374986
    18173 1kzo:B (2.2) BS02 ZN 2.5.1.58 GO:0003824 ... Q02293 12374986
    18174 1kzo:B (2.2) BS03 FPP 2.5.1.58 GO:0003824 ... Q02293 12374986 MOAD: Kd=5nM
    18175 1kzo:B (2.2) BS04 FAR 2.5.1.58 GO:0003824 ... Q02293 12374986
    18176 1kzp:B (2.1) BS01 peptide 2.5.1.58 GO:0003824 ... Q02293 12374986
    18177 1kzp:B (2.1) BS02 ZN 2.5.1.58 GO:0003824 ... Q02293 12374986
    18178 1kzp:B (2.1) BS03 FAR 2.5.1.58 GO:0003824 ... Q02293 12374986
    18179 1kzu:A (2.5) BS01 BCL ? GO:0005886 ... P26789 9159480
    18180 1kzu:A (2.5) BS02 BCL ? GO:0005886 ... P26789 9159480
    18181 1kzu:A (2.5) BS03 BCL ? GO:0005886 ... P26789 9159480
    18182 1kzu:B (2.5) BS01 BCL ? GO:0005886 ... P26790 9159480
    18183 1kzu:B (2.5) BS02 BCL ? GO:0005886 ... P26790 9159480
    18184 1kzu:B (2.5) BS03 BCL ? GO:0005886 ... P26790 9159480
    18185 1kzu:D (2.5) BS01 BCL ? GO:0005886 ... P26789 9159480
    18186 1kzu:D (2.5) BS02 BCL ? GO:0005886 ... P26789 9159480
    18187 1kzu:D (2.5) BS03 BCL ? GO:0005886 ... P26789 9159480
    18188 1kzu:D (2.5) BS04 BCL ? GO:0005886 ... P26789 9159480
    18189 1kzu:E (2.5) BS01 BCL ? GO:0005886 ... P26790 9159480
    18190 1kzu:E (2.5) BS02 BCL ? GO:0005886 ... P26790 9159480
    18191 1kzu:E (2.5) BS03 BCL ? GO:0005886 ... P26790 9159480
    18192 1kzu:G (2.5) BS01 BCL ? GO:0005886 ... P26789 9159480
    18193 1kzu:G (2.5) BS02 BCL ? GO:0005886 ... P26789 9159480
    18194 1kzu:G (2.5) BS03 BCL ? GO:0005886 ... P26789 9159480
    18195 1kzu:G (2.5) BS04 BCL ? GO:0005886 ... P26789 9159480
    18196 1kzu:H (2.5) BS01 BCL ? GO:0005886 ... P26790 9159480
    18197 1kzu:H (2.5) BS02 BCL ? GO:0005886 ... P26790 9159480
    18198 1kzu:H (2.5) BS03 BCL ? GO:0005886 ... P26790 9159480
    18199 1kzy:A (2.5) BS01 ZN ? GO:0000976 ... P04637 11877378
    18200 1kzy:B (2.5) BS01 ZN ? GO:0000976 ... P04637 11877378

    Reference:
  • Chengxin Zhang, Xi Zhang, Peter L Freddolino, and Yang Zhang. BioLiP2: an updated structure database for biologically relevent ligand-protein interactions, Nucleic Acids Research, gkad630 (2023).
  • Jianyi Yang, Ambrish Roy, and Yang Zhang. BioLiP: a semi-manually curated database for biologically relevant ligand-protein interactions, Nucleic Acids Research, 41: D1096-D1103 (2013) (download the PDF file).
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