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BioLiP
Download all results in tab-seperated text for 594392 receptor-ligand interactions, whose format is explained at readme.txt.
  • Hover over PDB to view the title of the structure. Click PDB to view the structure at the RCSB PDB database. Resolution -1.00 means the resolution is unavailable, e.g., for NMR structures.
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  • Hover over Ligand to view the full ligand name. Click Ligand to view the 2D diagram and other detail information of the ligand.
  • Hover over EC number to view the full name of enzymatic activity.
  • Hover over GO terms to view all GO terms.
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    # PDB
    (Resolution Å)
    Site # Ligand EC number GO terms UniProt PubMed Binding
    affinity
    59401 2otl:O (2.7) BS01 rna ? GO:0003723 ... P12733 17321546
    59402 2otl:P (2.7) BS01 rna ? GO:0003723 ... P14119 17321546
    59403 2otl:Q (2.7) BS01 rna ? GO:0003735 ... P12734 17321546
    59404 2otl:Q (2.7) BS02 rna ? GO:0003735 ... P12734 17321546
    59405 2otl:R (2.7) BS01 rna ? GO:0002181 ... P10970 17321546
    59406 2otl:S (2.7) BS01 rna ? GO:0000027 ... P12732 17321546
    59407 2otl:T (2.7) BS01 rna ? GO:0002181 ... P10972 17321546
    59408 2otl:T (2.7) BS02 MG ? GO:0002181 ... P10972 17321546
    59409 2otl:U (2.7) BS01 rna ? GO:0003735 ... P14116 17321546
    59410 2otl:V (2.7) BS01 rna ? GO:0003735 ... P10971 17321546
    59411 2otl:W (2.7) BS01 rna ? GO:0000463 ... P14121 17321546
    59412 2otl:W (2.7) BS02 rna ? GO:0000463 ... P14121 17321546
    59413 2otl:X (2.7) BS01 rna ? GO:0002181 ... P18138 17321546
    59414 2otl:Y (2.7) BS01 rna ? GO:0000166 ... P12736 17321546
    59415 2otl:Y (2.7) BS02 MG ? GO:0000166 ... P12736 17321546
    59416 2otl:Z (2.7) BS01 rna ? GO:0003735 ... P60619 17321546
    59417 2otv:A (1.56) BS01 CA 3.4.21.4 GO:0004175 ... P00760 N/A
    59418 2otv:A (1.56) BS02 NCA 3.4.21.4 GO:0004175 ... P00760 N/A
    59419 2oty:X (1.83) BS01 YAN 3.2.1.17 GO:0003796 ... P00720 17599350 MOAD: Kd=21uM
    59420 2otz:X (2.07) BS01 1MR 3.2.1.17 GO:0003796 ... P00720 17599350 MOAD: Kd=357uM
    59421 2ou0:X (1.94) BS01 MR3 3.2.1.17 GO:0003796 ... P00720 17599350 MOAD: Kd=592uM
    59422 2ou8:A (1.8) BS01 MTN 3.2.1.17 GO:0003796 ... P00720 17473014
    59423 2ou9:A (1.55) BS01 MTN 3.2.1.17 GO:0003796 ... P00720 17473014
    59424 2oub:A (2.75) BS01 2TN 3.1.1.4 GO:0004623 ... P59071 N/A
    59425 2oui:A (1.77) BS01 ZN 1.1.1.80 GO:0005737 ... P35630 18260103
    59426 2oui:B (1.77) BS01 ZN 1.1.1.80 GO:0005737 ... P35630 18260103
    59427 2oui:C (1.77) BS01 ZN 1.1.1.80 GO:0005737 ... P35630 18260103
    59428 2oui:D (1.77) BS01 ZN 1.1.1.80 GO:0005737 ... P35630 18260103
    59429 2oux:A (2.16) BS01 MG ? GO:0015095 ... Q830V1 19787770
    59430 2oux:A (2.16) BS02 MG ? GO:0015095 ... Q830V1 19787770
    59431 2oux:A (2.16) BS03 MG ? GO:0015095 ... Q830V1 19787770
    59432 2oux:A (2.16) BS04 MG ? GO:0015095 ... Q830V1 19787770
    59433 2oux:B (2.16) BS01 MG ? GO:0015095 ... Q830V1 19787770
    59434 2oux:B (2.16) BS02 MG ? GO:0015095 ... Q830V1 19787770
    59435 2oux:B (2.16) BS03 MG ? GO:0015095 ... Q830V1 19787770
    59436 2oux:B (2.16) BS04 MG ? GO:0015095 ... Q830V1 19787770
    59437 2ov0:A (0.75) BS01 CU ? GO:0005507 ... P22364 N/A
    59438 2ov2:A (2.1) BS01 MG 3.6.5.2 GO:0003924 ... P60763 N/A
    59439 2ov2:B (2.1) BS01 MG 3.6.5.2 GO:0003924 ... P60763 N/A
    59440 2ov2:C (2.1) BS01 MG 3.6.5.2 GO:0003924 ... P60763 N/A
    59441 2ov2:D (2.1) BS01 MG 3.6.5.2 GO:0003924 ... P60763 N/A
    59442 2ov2:E (2.1) BS01 MG 3.6.5.2 GO:0003924 ... P60763 N/A
    59443 2ov2:F (2.1) BS01 MG 3.6.5.2 GO:0003924 ... P60763 N/A
    59444 2ov2:G (2.1) BS01 MG 3.6.5.2 GO:0003924 ... P60763 N/A
    59445 2ov2:H (2.1) BS01 MG 3.6.5.2 GO:0003924 ... P60763 N/A
    59446 2ov4:A (2.5) BS01 AQP 6.1.1.2 GO:0000166 ... P00953 17428498 MOAD: Kd=18uM
    PDBbind: -logKd/Ki=4.74, Kd=18uM
    59447 2ov4:A (2.5) BS02 ANL 6.1.1.2 GO:0000166 ... P00953 17428498
    59448 2ovb:A (2.61) BS01 SO4 2.8.2.36 GO:0005737 ... Q8KLM3 17329243
    59449 2ovb:A (2.61) BS02 SO4 2.8.2.36 GO:0005737 ... Q8KLM3 17329243
    59450 2ovb:A (2.61) BS03 SO4 2.8.2.36 GO:0005737 ... Q8KLM3 17329243
    59451 2ovd:A (1.8) BS01 DAO ? GO:0001848 ... P07360 17452033
    59452 2ovf:A (2.95) BS01 A3P 2.8.2.36 GO:0005737 ... Q8KLM3 17329243
    59453 2ovx:A (2.0) BS01 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59454 2ovx:A (2.0) BS02 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59455 2ovx:A (2.0) BS03 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59456 2ovx:A (2.0) BS04 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59457 2ovx:A (2.0) BS05 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59458 2ovx:A (2.0) BS06 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59459 2ovx:A (2.0) BS07 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59460 2ovx:A (2.0) BS08 4MR 3.4.24.35 GO:0004222 ... P14780 17599356 PDBbind: -logKd/Ki=8.70, IC50=2nM
    59461 2ovx:B (2.0) BS01 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59462 2ovx:B (2.0) BS02 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59463 2ovx:B (2.0) BS03 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59464 2ovx:B (2.0) BS04 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59465 2ovx:B (2.0) BS05 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59466 2ovx:B (2.0) BS06 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59467 2ovx:B (2.0) BS07 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59468 2ovx:B (2.0) BS08 4MR 3.4.24.35 GO:0004222 ... P14780 17599356
    59469 2ovz:A (2.0) BS01 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59470 2ovz:A (2.0) BS02 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59471 2ovz:A (2.0) BS03 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59472 2ovz:A (2.0) BS04 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59473 2ovz:A (2.0) BS05 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59474 2ovz:A (2.0) BS06 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59475 2ovz:A (2.0) BS07 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59476 2ovz:A (2.0) BS08 5MR 3.4.24.35 GO:0004222 ... P14780 17599356 PDBbind: -logKd/Ki=7.89, Ki=13nM
    59477 2ovz:B (2.0) BS01 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59478 2ovz:B (2.0) BS02 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59479 2ovz:B (2.0) BS03 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59480 2ovz:B (2.0) BS04 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59481 2ovz:B (2.0) BS05 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59482 2ovz:B (2.0) BS06 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59483 2ovz:B (2.0) BS07 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59484 2ovz:B (2.0) BS08 5MR 3.4.24.35 GO:0004222 ... P14780 17599356
    59485 2ow0:A (2.0) BS01 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59486 2ow0:A (2.0) BS02 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59487 2ow0:A (2.0) BS03 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59488 2ow0:A (2.0) BS04 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59489 2ow0:A (2.0) BS05 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59490 2ow0:A (2.0) BS06 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59491 2ow0:A (2.0) BS07 6MR 3.4.24.35 GO:0004222 ... P14780 17599356 PDBbind: -logKd/Ki=6.70, IC50=201nM
    59492 2ow0:B (2.0) BS01 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59493 2ow0:B (2.0) BS02 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59494 2ow0:B (2.0) BS03 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59495 2ow0:B (2.0) BS04 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59496 2ow0:B (2.0) BS05 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59497 2ow0:B (2.0) BS06 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59498 2ow0:B (2.0) BS07 6MR 3.4.24.35 GO:0004222 ... P14780 17599356
    59499 2ow1:A (2.2) BS01 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59500 2ow1:A (2.2) BS02 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59501 2ow1:A (2.2) BS03 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59502 2ow1:A (2.2) BS04 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59503 2ow1:A (2.2) BS05 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59504 2ow1:A (2.2) BS06 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59505 2ow1:A (2.2) BS07 7MR 3.4.24.35 GO:0004222 ... P14780 17599356 PDBbind: -logKd/Ki=9.00, IC50=1nM
    59506 2ow1:B (2.2) BS01 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59507 2ow1:B (2.2) BS02 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59508 2ow1:B (2.2) BS03 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59509 2ow1:B (2.2) BS04 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59510 2ow1:B (2.2) BS05 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59511 2ow1:B (2.2) BS06 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59512 2ow1:B (2.2) BS07 7MR 3.4.24.35 GO:0004222 ... P14780 17599356
    59513 2ow2:A (2.9) BS01 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59514 2ow2:A (2.9) BS02 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59515 2ow2:A (2.9) BS03 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59516 2ow2:A (2.9) BS04 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59517 2ow2:A (2.9) BS05 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59518 2ow2:A (2.9) BS06 8MR 3.4.24.35 GO:0004222 ... P14780 17599356 PDBbind: -logKd/Ki=8.22, IC50=6nM
    59519 2ow2:B (2.9) BS01 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59520 2ow2:B (2.9) BS02 ZN 3.4.24.35 GO:0004222 ... P14780 17599356
    59521 2ow2:B (2.9) BS03 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59522 2ow2:B (2.9) BS04 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59523 2ow2:B (2.9) BS05 CA 3.4.24.35 GO:0004222 ... P14780 17599356
    59524 2ow2:B (2.9) BS06 8MR 3.4.24.35 GO:0004222 ... P14780 17599356
    59525 2ow6:A (1.19) BS01 ZN 3.2.1.114 GO:0000139 ... Q24451 18076078
    59526 2ow6:A (1.19) BS02 NK1 3.2.1.114 GO:0000139 ... Q24451 18076078 MOAD: ic50=2mM
    PDBbind: -logKd/Ki=2.70, IC50=2mM
    59527 2ow7:A (1.77) BS01 ZN 3.2.1.114 GO:0000139 ... Q24451 18076078
    59528 2ow7:A (1.77) BS02 NK2 3.2.1.114 GO:0000139 ... Q24451 18076078 MOAD: ic50=14uM
    PDBbind: -logKd/Ki=4.85, IC50=0.014mM
    59529 2ow9:A (1.74) BS01 ZN 3.4.24.- GO:0004222 ... P45452 17623656
    59530 2ow9:A (1.74) BS02 ZN 3.4.24.- GO:0004222 ... P45452 17623656
    59531 2ow9:A (1.74) BS03 CA 3.4.24.- GO:0004222 ... P45452 17623656
    59532 2ow9:A (1.74) BS04 CA 3.4.24.- GO:0004222 ... P45452 17623656
    59533 2ow9:A (1.74) BS05 CA 3.4.24.- GO:0004222 ... P45452 17623656
    59534 2ow9:A (1.74) BS06 SP6 3.4.24.- GO:0004222 ... P45452 17623656 MOAD: ic50=30nM
    PDBbind: -logKd/Ki=7.52, IC50=30nM
    BindingDB: IC50=30nM, Ki=64nM
    59535 2ow9:B (1.74) BS01 ZN 3.4.24.- GO:0004222 ... P45452 17623656
    59536 2ow9:B (1.74) BS02 ZN 3.4.24.- GO:0004222 ... P45452 17623656
    59537 2ow9:B (1.74) BS03 CA 3.4.24.- GO:0004222 ... P45452 17623656
    59538 2ow9:B (1.74) BS04 CA 3.4.24.- GO:0004222 ... P45452 17623656
    59539 2ow9:B (1.74) BS05 CA 3.4.24.- GO:0004222 ... P45452 17623656
    59540 2ow9:B (1.74) BS06 SP6 3.4.24.- GO:0004222 ... P45452 17623656 MOAD: ic50=30nM
    BindingDB: IC50=30nM, Ki=64nM
    59541 2owc:A (2.05) BS01 GLC 2.4.1.25 GO:0004134 ... O87172 17420245
    59542 2owc:A (2.05) BS02 AC1 2.4.1.25 GO:0004134 ... O87172 17420245
    59543 2owl:A (2.4) BS01 CA ? GO:0000018 ... P36767 17308310
    59544 2owl:B (2.4) BS01 CA ? GO:0000018 ... P36767 17308310
    59545 2owo:A (2.3) BS01 dna 6.5.1.2 GO:0003677 ... P15042 17466627
    59546 2owo:A (2.3) BS02 dna 6.5.1.2 GO:0003677 ... P15042 17466627
    59547 2owo:A (2.3) BS03 dna 6.5.1.2 GO:0003677 ... P15042 17466627
    59548 2owo:A (2.3) BS04 AMP 6.5.1.2 GO:0003677 ... P15042 17466627
    59549 2owo:A (2.3) BS05 ZN 6.5.1.2 GO:0003677 ... P15042 17466627
    59550 2ows:A (1.49) BS01 FE ? GO:0030288 ... Q7VXW9 N/A
    59551 2ows:A (1.49) BS02 OXL ? GO:0030288 ... Q7VXW9 N/A
    59552 2owt:A (2.4) BS01 FE ? GO:0030288 ... Q7VXW9 N/A
    59553 2owt:A (2.4) BS02 CO3 ? GO:0030288 ... Q7VXW9 N/A
    59554 2oww:A (2.2) BS01 GLC 2.4.1.25 GO:0004134 ... O87172 17420245
    59555 2oww:A (2.2) BS02 AC1 2.4.1.25 GO:0004134 ... O87172 17420245
    59556 2oww:A (2.2) BS03 G4D 2.4.1.25 GO:0004134 ... O87172 17420245
    59557 2owz:A (2.18) BS01 F6P 3.1.3.11 GO:0000166 ... P0A993 17314096
    59558 2ox3:A (2.18) BS01 F6P 3.1.3.11 GO:0000166 ... P0A993 17314096
    59559 2ox3:A (2.18) BS02 PEP 3.1.3.11 GO:0000166 ... P0A993 17314096
    59560 2ox6:A (1.7) BS01 MG ? GO:0003674 ... Q8EAP9 N/A
    59561 2oxb:A (2.6) BS01 FRU 3.2.1.26 GO:0004553 ... Q43866 17963237
    59562 2oxd:A (2.3) BS01 K32 2.7.11.1 GO:0004672 ... P28523 17768728 PDBbind: -logKd/Ki=6.82, Ki=0.15uM
    59563 2oxe:A (2.8) BS01 CA 3.1.1.26
    3.1.1.3
    GO:0004620 ... P54317 18702514
    59564 2oxe:B (2.8) BS01 MAN 3.1.1.26
    3.1.1.3
    GO:0004620 ... P54317 18702514
    59565 2oxe:B (2.8) BS02 CA 3.1.1.26
    3.1.1.3
    GO:0004620 ... P54317 18702514
    59566 2oxi:A (2.1) BS01 CU 1.1.1.1 GO:0004022 ... P00327 15299812
    59567 2oxi:A (2.1) BS02 ZN 1.1.1.1 GO:0004022 ... P00327 15299812
    59568 2oxi:A (2.1) BS03 NAD 1.1.1.1 GO:0004022 ... P00327 15299812
    59569 2oxi:A (2.1) BS04 DMS 1.1.1.1 GO:0004022 ... P00327 15299812
    59570 2oxi:B (2.1) BS01 CU 1.1.1.1 GO:0004022 ... P00327 15299812
    59571 2oxi:B (2.1) BS02 ZN 1.1.1.1 GO:0004022 ... P00327 15299812
    59572 2oxi:B (2.1) BS03 NAD 1.1.1.1 GO:0004022 ... P00327 15299812
    59573 2oxi:B (2.1) BS04 DMS 1.1.1.1 GO:0004022 ... P00327 15299812
    59574 2oxn:A (1.7) BS01 OAN 3.2.1.52 GO:0004553 ... Q9KU37 17439950 MOAD: Ki=0.036uM
    PDBbind: -logKd/Ki=7.44, Ki=36nM
    59575 2oxs:A (1.32) BS01 CA 3.4.21.4 GO:0004175 ... P00760 N/A
    59576 2oxu:A (1.24) BS01 ZN 3.4.24.65 GO:0004222 ... P39900 17096442
    59577 2oxu:A (1.24) BS02 ZN 3.4.24.65 GO:0004222 ... P39900 17096442
    59578 2oxu:A (1.24) BS03 CA 3.4.24.65 GO:0004222 ... P39900 17096442
    59579 2oxu:A (1.24) BS04 CA 3.4.24.65 GO:0004222 ... P39900 17096442
    59580 2oxu:A (1.24) BS05 CA 3.4.24.65 GO:0004222 ... P39900 17096442
    59581 2oxw:A (1.15) BS01 peptide 3.4.24.65 GO:0004222 ... P39900 17096442
    59582 2oxw:A (1.15) BS02 ZN 3.4.24.65 GO:0004222 ... P39900 17096442
    59583 2oxw:A (1.15) BS03 ZN 3.4.24.65 GO:0004222 ... P39900 17096442
    59584 2oxw:A (1.15) BS04 CA 3.4.24.65 GO:0004222 ... P39900 17096442
    59585 2oxw:A (1.15) BS05 CA 3.4.24.65 GO:0004222 ... P39900 17096442
    59586 2oxw:A (1.15) BS06 CA 3.4.24.65 GO:0004222 ... P39900 17096442
    59587 2oxx:A (2.3) BS01 K22 2.7.11.1 GO:0004672 ... P28523 17768728 PDBbind: -logKd/Ki=6.70, Ki=0.20uM
    59588 2oxy:A (1.812) BS01 K17 2.7.11.1 GO:0004672 ... P28523 17768728 PDBbind: -logKd/Ki=6.52, Ki=0.30uM
    59589 2oxy:B (1.812) BS01 K17 2.7.11.1 GO:0004672 ... P28523 17768728
    59590 2oxz:A (1.9) BS01 peptide 3.4.24.65 GO:0004222 ... P39900 17096442
    59591 2oxz:A (1.9) BS02 peptide 3.4.24.65 GO:0004222 ... P39900 17096442
    59592 2oxz:A (1.9) BS03 ZN 3.4.24.65 GO:0004222 ... P39900 17096442
    59593 2oxz:A (1.9) BS04 ZN 3.4.24.65 GO:0004222 ... P39900 17096442
    59594 2oxz:A (1.9) BS05 CA 3.4.24.65 GO:0004222 ... P39900 17096442
    59595 2oxz:A (1.9) BS06 CA 3.4.24.65 GO:0004222 ... P39900 17096442
    59596 2oxz:A (1.9) BS07 CA 3.4.24.65 GO:0004222 ... P39900 17096442
    59597 2oy2:A (1.5) BS01 peptide 3.4.24.34 GO:0004222 ... P22894 17096442
    59598 2oy2:A (1.5) BS02 CA 3.4.24.34 GO:0004222 ... P22894 17096442
    59599 2oy2:A (1.5) BS03 CA 3.4.24.34 GO:0004222 ... P22894 17096442
    59600 2oy2:A (1.5) BS04 ZN 3.4.24.34 GO:0004222 ... P22894 17096442

    Reference:
  • Chengxin Zhang, Xi Zhang, Peter L Freddolino, and Yang Zhang. BioLiP2: an updated structure database for biologically relevent ligand-protein interactions, Nucleic Acids Research, gkad630 (2023).
  • Jianyi Yang, Ambrish Roy, and Yang Zhang. BioLiP: a semi-manually curated database for biologically relevant ligand-protein interactions, Nucleic Acids Research, 41: D1096-D1103 (2013) (download the PDF file).
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