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BioLiP
Download all results in tab-seperated text for 817662 receptor-ligand interactions, whose format is explained at readme.txt.
  • Hover over PDB to view the title of the structure. Click PDB to view the structure at the RCSB PDB database. Resolution -1.00 means the resolution is unavailable, e.g., for NMR structures.
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    # PDB
    (Resolution Å)
    Site # Ligand EC number GO terms UniProt PubMed Binding
    affinity
    43201 1rcn:E (2.32) BS01 dna 4.6.1.18 GO:0003676 ... P61823 8063789
    43202 1rco:B (2.3) BS01 XDP 4.1.1.39 GO:0000287 ... P00875 8955130
    43203 1rco:B (2.3) BS02 XDP 4.1.1.39 GO:0000287 ... P00875 8955130
    43204 1rco:E (2.3) BS01 XDP 4.1.1.39 GO:0000287 ... P00875 8955130
    43205 1rco:E (2.3) BS02 XDP 4.1.1.39 GO:0000287 ... P00875 8955130
    43206 1rco:H (2.3) BS01 XDP 4.1.1.39 GO:0000287 ... P00875 8955130
    43207 1rco:H (2.3) BS02 XDP 4.1.1.39 GO:0000287 ... P00875 8955130
    43208 1rco:K (2.3) BS01 XDP 4.1.1.39 GO:0000287 ... P00875 8955130
    43209 1rco:K (2.3) BS02 XDP 4.1.1.39 GO:0000287 ... P00875 8955130
    43210 1rco:L (2.3) BS01 XDP 4.1.1.39 GO:0000287 ... P00875 8955130
    43211 1rco:L (2.3) BS02 XDP 4.1.1.39 GO:0000287 ... P00875 8955130
    43212 1rco:O (2.3) BS01 XDP 4.1.1.39 GO:0000287 ... P00875 8955130
    43213 1rco:O (2.3) BS02 XDP 4.1.1.39 GO:0000287 ... P00875 8955130
    43214 1rco:R (2.3) BS01 XDP 4.1.1.39 GO:0000287 ... P00875 8955130
    43215 1rco:V (2.3) BS01 XDP 4.1.1.39 GO:0000287 ... P00875 8955130
    43216 1rco:V (2.3) BS02 XDP 4.1.1.39 GO:0000287 ... P00875 8955130
    43217 1rcp:A (2.0) BS01 HEM ? GO:0005506 ... P00147 8676382
    43218 1rcp:B (2.0) BS01 HEM ? GO:0005506 ... P00147 8676382
    43219 1rcq:A (1.45) BS01 PLP 5.1.1.1 GO:0003824 ... Q9HTQ2 14674749
    43220 1rcq:A (1.45) BS02 DLY 5.1.1.1 GO:0003824 ... Q9HTQ2 14674749
    43221 1rcs:A (-1.00) BS01 dna ? GO:0003677 ... P0A881 8176748
    43222 1rcs:A (-1.00) BS02 dna ? GO:0003677 ... P0A881 8176748
    43223 1rcs:A (-1.00) BS03 TRP ? GO:0003677 ... P0A881 8176748
    43224 1rcs:B (-1.00) BS01 dna ? GO:0003677 ... P0A881 8176748
    43225 1rcs:B (-1.00) BS02 dna ? GO:0003677 ... P0A881 8176748
    43226 1rcs:B (-1.00) BS03 TRP ? GO:0003677 ... P0A881 8176748
    43227 1rct:E (2.8) BS01 NOS 2.4.2.1 GO:0000255 ... P00491 14706628
    43228 1rcv:D (1.6) BS01 BV1 ? GO:0005515 ... P01556 15380181 MOAD: ic50=29uM
    43229 1rcv:E (1.6) BS01 BV1 ? GO:0005515 ... P01556 15380181 MOAD: ic50=29uM
    43230 1rcv:F (1.6) BS01 BV1 ? GO:0005515 ... P01556 15380181 MOAD: ic50=29uM
    43231 1rcv:F (1.6) BS02 BV1 ? GO:0005515 ... P01556 15380181 MOAD: ic50=29uM
    43232 1rcv:G (1.6) BS01 BV1 ? GO:0005515 ... P01556 15380181 MOAD: ic50=29uM
    43233 1rcv:H (1.6) BS01 BV1 ? GO:0005515 ... P01556 15380181 MOAD: ic50=29uM
    43234 1rcw:A (2.5) BS01 FE 1.3.3.- GO:0005576 ... O84616 15087448
    43235 1rcw:A (2.5) BS02 FE 1.3.3.- GO:0005576 ... O84616 15087448
    43236 1rcw:B (2.5) BS01 FE 1.3.3.- GO:0005576 ... O84616 15087448
    43237 1rcw:B (2.5) BS02 FE 1.3.3.- GO:0005576 ... O84616 15087448
    43238 1rcw:C (2.5) BS01 FE 1.3.3.- GO:0005576 ... O84616 15087448
    43239 1rcw:C (2.5) BS02 FE 1.3.3.- GO:0005576 ... O84616 15087448
    43240 1rcx:B (2.4) BS01 RUB 4.1.1.39 GO:0000287 ... P00875 9034362
    43241 1rcx:B (2.4) BS02 RUB 4.1.1.39 GO:0000287 ... P00875 9034362
    43242 1rcx:E (2.4) BS01 RUB 4.1.1.39 GO:0000287 ... P00875 9034362
    43243 1rcx:E (2.4) BS02 RUB 4.1.1.39 GO:0000287 ... P00875 9034362
    43244 1rcx:H (2.4) BS01 RUB 4.1.1.39 GO:0000287 ... P00875 9034362
    43245 1rcx:H (2.4) BS02 RUB 4.1.1.39 GO:0000287 ... P00875 9034362
    43246 1rcx:K (2.4) BS01 RUB 4.1.1.39 GO:0000287 ... P00875 9034362
    43247 1rcx:K (2.4) BS02 RUB 4.1.1.39 GO:0000287 ... P00875 9034362
    43248 1rcx:L (2.4) BS01 RUB 4.1.1.39 GO:0000287 ... P00875 9034362
    43249 1rcx:L (2.4) BS02 RUB 4.1.1.39 GO:0000287 ... P00875 9034362
    43250 1rcx:O (2.4) BS01 RUB 4.1.1.39 GO:0000287 ... P00875 9034362
    43251 1rcx:O (2.4) BS02 RUB 4.1.1.39 GO:0000287 ... P00875 9034362
    43252 1rcx:R (2.4) BS01 RUB 4.1.1.39 GO:0000287 ... P00875 9034362
    43253 1rcx:R (2.4) BS02 RUB 4.1.1.39 GO:0000287 ... P00875 9034362
    43254 1rcx:V (2.4) BS01 RUB 4.1.1.39 GO:0000287 ... P00875 9034362
    43255 1rcx:V (2.4) BS02 RUB 4.1.1.39 GO:0000287 ... P00875 9034362
    43256 1rcy:A (1.9) BS01 CU ? GO:0005507 ... P0C918 8947572
    43257 1rd9:D (1.44) BS01 BV2 ? GO:0005515 ... P01556 15380181 MOAD: ic50=13uM
    43258 1rd9:D (1.44) BS02 BV2 ? GO:0005515 ... P01556 15380181 MOAD: ic50=13uM
    43259 1rd9:E (1.44) BS01 BV2 ? GO:0005515 ... P01556 15380181 MOAD: ic50=13uM
    43260 1rd9:F (1.44) BS01 BV2 ? GO:0005515 ... P01556 15380181 MOAD: ic50=13uM
    43261 1rd9:F (1.44) BS02 BV2 ? GO:0005515 ... P01556 15380181 MOAD: ic50=13uM
    43262 1rd9:G (1.44) BS01 BV2 ? GO:0005515 ... P01556 15380181 MOAD: ic50=13uM
    43263 1rd9:H (1.44) BS01 BV2 ? GO:0005515 ... P01556 15380181 MOAD: ic50=13uM
    43264 1rdd:A (2.8) BS01 MG 3.1.26.4 GO:0000287 ... P0A7Y4 8108376
    43265 1rdf:A (2.8) BS01 MG 3.11.1.1 GO:0000287 ... O31156 15005616
    43266 1rdf:A (2.8) BS02 ESA 3.11.1.1 GO:0000287 ... O31156 15005616
    43267 1rdf:B (2.8) BS01 MG 3.11.1.1 GO:0000287 ... O31156 15005616
    43268 1rdf:B (2.8) BS02 ESA 3.11.1.1 GO:0000287 ... O31156 15005616
    43269 1rdf:C (2.8) BS01 MG 3.11.1.1 GO:0000287 ... O31156 15005616
    43270 1rdf:C (2.8) BS02 ESA 3.11.1.1 GO:0000287 ... O31156 15005616
    43271 1rdf:D (2.8) BS01 MG 3.11.1.1 GO:0000287 ... O31156 15005616
    43272 1rdf:D (2.8) BS02 ESA 3.11.1.1 GO:0000287 ... O31156 15005616
    43273 1rdf:E (2.8) BS01 MG 3.11.1.1 GO:0000287 ... O31156 15005616
    43274 1rdf:E (2.8) BS02 ESA 3.11.1.1 GO:0000287 ... O31156 15005616
    43275 1rdf:F (2.8) BS01 MG 3.11.1.1 GO:0000287 ... O31156 15005616
    43276 1rdf:F (2.8) BS02 ESA 3.11.1.1 GO:0000287 ... O31156 15005616
    43277 1rdg:A (1.4) BS01 FE ? GO:0005506 ... P00270 3441010
    43278 1rdp:D (1.35) BS01 BV3 ? GO:0005515 ... P01556 15380181 MOAD: ic50=9uM
    43279 1rdp:E (1.35) BS01 BV3 ? GO:0005515 ... P01556 15380181 MOAD: ic50=9uM
    43280 1rdp:F (1.35) BS01 BV3 ? GO:0005515 ... P01556 15380181 MOAD: ic50=9uM
    43281 1rdp:G (1.35) BS01 BV3 ? GO:0005515 ... P01556 15380181 MOAD: ic50=9uM
    43282 1rdp:H (1.35) BS01 BV3 ? GO:0005515 ... P01556 15380181 MOAD: ic50=9uM
    43283 1rdq:E (1.26) BS01 peptide 2.7.11.11 GO:0000122 ... P05132 14757059
    43284 1rdq:E (1.26) BS02 MG 2.7.11.11 GO:0000122 ... P05132 14757059
    43285 1rdq:E (1.26) BS03 ADP 2.7.11.11 GO:0000122 ... P05132 14757059
    43286 1rdq:E (1.26) BS04 ATP 2.7.11.11 GO:0000122 ... P05132 14757059
    43287 1rdr:A (2.4) BS01 CA 2.7.7.48
    3.4.22.28
    3.4.22.29
    3.6.1.15
    GO:0003723 ... P03300 9309225
    43288 1rds:A (1.8) BS01 GPC 4.6.1.24 GO:0003723 ... P00653 8218254
    43289 1rdt:A (2.4) BS01 peptide ? GO:0003677 ... P19793 15056000
    43290 1rdt:A (2.4) BS02 L79 ? GO:0003677 ... P19793 15056000 MOAD: Ki=6nM
    PDBbind: -logKd/Ki=6.70, Ki=200nM
    43291 1rdt:D (2.4) BS01 peptide ? GO:0003677 ... P37231 15056000
    43292 1rdt:D (2.4) BS02 570 ? GO:0003677 ... P37231 15056000 BindingDB: Ki=1.1nM, EC50=0.338844nM, IC50=25nM
    43293 1rdv:A (2.0) BS01 FE ? GO:0005506 ... P15412 10089348
    43294 1rdw:X (2.3) BS01 ATP 3.6.4.- GO:0000287 ... P68135 15099571
    43295 1rdw:X (2.3) BS02 LAR 3.6.4.- GO:0000287 ... P68135 15099571
    43296 1rdx:A (2.75) BS01 F6P 3.1.3.11 GO:0005737 ... P00636 8844845
    43297 1rdx:B (2.75) BS01 F6P 3.1.3.11 GO:0005737 ... P00636 8844845
    43298 1rdy:A (2.2) BS01 F6P 3.1.3.11 GO:0005737 ... P00636 8844845
    43299 1rdy:A (2.2) BS02 AMP 3.1.3.11 GO:0005737 ... P00636 8844845 BindingDB: IC50=1300nM
    43300 1rdy:B (2.2) BS01 F6P 3.1.3.11 GO:0005737 ... P00636 8844845
    43301 1rdy:B (2.2) BS02 AMP 3.1.3.11 GO:0005737 ... P00636 8844845 BindingDB: IC50=1300nM
    43302 1rdz:A (2.05) BS01 F6P 3.1.3.11 GO:0005737 ... P00636 8844845
    43303 1rdz:A (2.05) BS02 AMP 3.1.3.11 GO:0005737 ... P00636 8844845 BindingDB: IC50=1300nM
    43304 1rdz:B (2.05) BS01 F6P 3.1.3.11 GO:0005737 ... P00636 8844845
    43305 1rdz:B (2.05) BS02 AMP 3.1.3.11 GO:0005737 ... P00636 8844845 BindingDB: IC50=1300nM
    43306 1re0:A (2.4) BS01 GDP 3.6.5.2 GO:0000139 ... P84077 14690595
    43307 1re0:A (2.4) BS02 AFB 3.6.5.2 GO:0000139 ... P84077 14690595
    43308 1re1:A (2.5) BS01 NA3 3.4.22.56 GO:0004197 ... P42574 15115390 PDBbind: -logKd/Ki=5.21, IC50=6117nM
    BindingDB: IC50=6000nM
    43309 1re1:B (2.5) BS01 NA3 3.4.22.56 GO:0004197 ... P42574 15115390 PDBbind: -logKd/Ki=5.21, IC50=6117nM
    BindingDB: IC50=6000nM
    43310 1re2:A (2.3) BS01 GAL 3.2.1.17 GO:0003796 ... P61626 9888793
    43311 1re3:B (2.45) BS01 CA ? GO:0005102 ... P02675 14992584
    43312 1re3:C (2.45) BS01 peptide ? GO:0005102 ... P02679 14992584
    43313 1re3:C (2.45) BS02 CA ? GO:0005102 ... P02679 14992584
    43314 1re3:C (2.45) BS03 CA ? GO:0005102 ... P02679 14992584
    43315 1re3:E (2.45) BS01 CA ? GO:0005102 ... P02675 14992584
    43316 1re3:F (2.45) BS01 peptide ? GO:0005102 ... P02679 14992584
    43317 1re3:F (2.45) BS02 CA ? GO:0005102 ... P02679 14992584
    43318 1re4:B (2.7) BS01 CA ? GO:0005102 ... P02675 14992584
    43319 1re4:B (2.7) BS02 CA ? GO:0005102 ... P02675 14992584
    43320 1re4:C (2.7) BS01 CA ? GO:0005102 ... P02679 14992584
    43321 1re4:C (2.7) BS02 CA ? GO:0005102 ... P02679 14992584
    43322 1re4:E (2.7) BS01 CA ? GO:0005102 ... P02675 14992584
    43323 1re4:F (2.7) BS01 CA ? GO:0005102 ... P02679 14992584
    43324 1re4:F (2.7) BS02 CA ? GO:0005102 ... P02679 14992584
    43325 1re8:A (2.1) BS01 BD2 2.7.11.11 GO:0000122 ... P05132 14705934 MOAD: Ki=0.3nM
    PDBbind: -logKd/Ki=9.52, Ki=0.3nM
    43326 1re9:A (1.45) BS01 HEM 1.14.15.1 GO:0004497 ... P00183 15522298
    43327 1re9:A (1.45) BS02 DSO 1.14.15.1 GO:0004497 ... P00183 15522298 MOAD: Kd~0.02uM
    43328 1rea:A (2.7) BS01 ADP ? GO:0000725 ... P0A7G6 1731253
    43329 1rec:A (1.9) BS01 CA ? GO:0001750 ... P21457 8242744
    43330 1red:B (1.6) BS01 C5X 3.2.1.8 GO:0004553 ... P36217 8755744
    43331 1ree:B (1.6) BS01 07E 3.2.1.8 GO:0004553 ... P36217 8755744
    43332 1ref:B (1.8) BS01 C3X 3.2.1.8 GO:0004553 ... P36217 8755744
    43333 1rej:A (2.2) BS01 B1L 2.7.11.11 GO:0000122 ... P05132 14705934 MOAD: Ki=5nM
    PDBbind: -logKd/Ki=8.30, Ki=5nM
    43334 1rek:A (2.3) BS01 B8L 2.7.11.11 GO:0000122 ... P05132 14705934 MOAD: Ki=200nM
    PDBbind: -logKd/Ki=6.70, Ki=200nM
    43335 1rek:A (2.3) BS02 PTL 2.7.11.11 GO:0000122 ... P05132 14705934
    43336 1rem:A (2.1) BS01 BMA 3.2.1.17 GO:0003796 ... P61626 9757098
    43337 1reo:A (2.31) BS01 FAD 1.4.3.2 GO:0001716 ... Q6STF1 15103157
    43338 1rep:C (2.6) BS01 dna ? GO:0003677 ... P03856 10469640
    43339 1rep:C (2.6) BS02 dna ? GO:0003677 ... P03856 10469640
    43340 1rep:C (2.6) BS03 MG ? GO:0003677 ... P03856 10469640
    43341 1req:A (2.0) BS01 B12 5.4.99.2 GO:0003824 ... P11653 8805541
    43342 1req:A (2.0) BS02 DCA 5.4.99.2 GO:0003824 ... P11653 8805541
    43343 1req:C (2.0) BS01 B12 5.4.99.2 GO:0003824 ... P11653 8805541
    43344 1req:C (2.0) BS02 DCA 5.4.99.2 GO:0003824 ... P11653 8805541
    43345 1req:D (2.0) BS01 DCA 5.4.99.2 GO:0003824 ... P11652 8805541
    43346 1rev:A (2.6) BS01 TB9 2.7.7.-
    2.7.7.49
    2.7.7.7
    3.1.-.-
    3.1.13.2
    3.1.26.13
    3.4.23.16
    GO:0003676 ... P04585 8535785 PDBbind: -logKd/Ki=7.48, Ki=33nM
    43347 1rez:A (1.7) BS01 GAL 3.2.1.17 GO:0003796 ... P61626 8885835
    43348 1rf0:B (2.81) BS01 CA ? GO:0005102 ... P02675 14992585
    43349 1rf0:B (2.81) BS02 CA ? GO:0005102 ... P02675 14992585
    43350 1rf0:C (2.81) BS01 CA ? GO:0005102 ... P02679 14992585
    43351 1rf0:E (2.81) BS01 CA ? GO:0005102 ... P02675 14992585
    43352 1rf0:E (2.81) BS02 CA ? GO:0005102 ... P02675 14992585
    43353 1rf0:F (2.81) BS01 CA ? GO:0005102 ... P02679 14992585
    43354 1rf1:B (2.53) BS01 peptide ? GO:0005102 ... P02675 14992585
    43355 1rf1:B (2.53) BS02 CA ? GO:0005102 ... P02675 14992585
    43356 1rf1:C (2.53) BS01 peptide ? GO:0005102 ... P02679 14992585
    43357 1rf1:C (2.53) BS02 CA ? GO:0005102 ... P02679 14992585
    43358 1rf1:E (2.53) BS01 peptide ? GO:0005102 ... P02675 14992585
    43359 1rf1:E (2.53) BS02 CA ? GO:0005102 ... P02675 14992585
    43360 1rf1:F (2.53) BS01 peptide ? GO:0005102 ... P02679 14992585
    43361 1rf1:F (2.53) BS02 CA ? GO:0005102 ... P02679 14992585
    43362 1rf2:D (1.35) BS01 BV4 ? GO:0005515 ... P01556 15380181 MOAD: ic50=17uM
    43363 1rf2:E (1.35) BS01 BV4 ? GO:0005515 ... P01556 15380181 MOAD: ic50=17uM
    43364 1rf2:F (1.35) BS01 BV4 ? GO:0005515 ... P01556 15380181 MOAD: ic50=17uM
    43365 1rf2:G (1.35) BS01 BV4 ? GO:0005515 ... P01556 15380181 MOAD: ic50=17uM
    43366 1rf2:H (1.35) BS01 BV4 ? GO:0005515 ... P01556 15380181 MOAD: ic50=17uM
    43367 1rf3:A (3.5) BS01 peptide 2.3.2.27 GO:0001817 ... Q13114 14517219
    43368 1rf4:A (2.2) BS01 SPQ 2.5.1.19 GO:0003824 ... Q9S400 14763973
    43369 1rf4:B (2.2) BS01 SPQ 2.5.1.19 GO:0003824 ... Q9S400 14763973
    43370 1rf4:C (2.2) BS01 SPQ 2.5.1.19 GO:0003824 ... Q9S400 14763973
    43371 1rf4:D (2.2) BS01 SPQ 2.5.1.19 GO:0003824 ... Q9S400 14763973
    43372 1rf6:A (1.9) BS01 GPJ 2.5.1.19 GO:0003824 ... Q9S400 14763973
    43373 1rf6:A (1.9) BS02 S3P 2.5.1.19 GO:0003824 ... Q9S400 14763973
    43374 1rf6:B (1.9) BS01 GPJ 2.5.1.19 GO:0003824 ... Q9S400 14763973
    43375 1rf6:B (1.9) BS02 S3P 2.5.1.19 GO:0003824 ... Q9S400 14763973
    43376 1rf6:C (1.9) BS01 GPJ 2.5.1.19 GO:0003824 ... Q9S400 14763973
    43377 1rf6:C (1.9) BS02 S3P 2.5.1.19 GO:0003824 ... Q9S400 14763973
    43378 1rf6:D (1.9) BS01 GPJ 2.5.1.19 GO:0003824 ... Q9S400 14763973
    43379 1rf6:D (1.9) BS02 S3P 2.5.1.19 GO:0003824 ... Q9S400 14763973
    43380 1rf7:A (1.8) BS01 DHF 1.5.1.3 GO:0004146 ... P0ABQ4 9012674
    43381 1rf7:A (1.8) BS02 DHF 1.5.1.3 GO:0004146 ... P0ABQ4 9012674
    43382 1rf8:A (-1.00) BS01 MTN ? GO:0000184 ... P07260 14675538
    43383 1rf8:A (-1.00) BS02 MTN ? GO:0000184 ... P07260 14675538
    43384 1rf8:A (-1.00) BS03 M7G ? GO:0000184 ... P07260 14675538
    43385 1rf9:A (1.8) BS01 HEM 1.14.15.1 GO:0004497 ... P00183 15522298
    43386 1rf9:A (1.8) BS02 DBR 1.14.15.1 GO:0004497 ... P00183 15522298
    43387 1rff:A (1.7) BS01 dna 3.1.4.- GO:0005634 ... Q9NUW8 14761185
    43388 1rff:A (1.7) BS02 peptide 3.1.4.- GO:0005634 ... Q9NUW8 14761185
    43389 1rff:A (1.7) BS03 VO4 3.1.4.- GO:0005634 ... Q9NUW8 14761185 BindingDB: IC50=4000nM
    43390 1rff:B (1.7) BS01 dna 3.1.4.- GO:0005634 ... Q9NUW8 14761185
    43391 1rff:B (1.7) BS02 peptide 3.1.4.- GO:0005634 ... Q9NUW8 14761185
    43392 1rff:B (1.7) BS03 VO4 3.1.4.- GO:0005634 ... Q9NUW8 14761185 BindingDB: IC50=4000nM
    43393 1rfg:E (2.9) BS01 GMP 2.4.2.1 GO:0000255 ... P00491 15983407
    43394 1rfi:A (2.2) BS01 dna 3.1.4.- GO:0005634 ... Q9NUW8 14761185
    43395 1rfi:A (2.2) BS02 peptide 3.1.4.- GO:0005634 ... Q9NUW8 14761185
    43396 1rfi:A (2.2) BS03 VO4 3.1.4.- GO:0005634 ... Q9NUW8 14761185 BindingDB: IC50=4000nM
    43397 1rfi:B (2.2) BS01 dna 3.1.4.- GO:0005634 ... Q9NUW8 14761185
    43398 1rfi:B (2.2) BS02 peptide 3.1.4.- GO:0005634 ... Q9NUW8 14761185
    43399 1rfi:B (2.2) BS03 VO4 3.1.4.- GO:0005634 ... Q9NUW8 14761185 BindingDB: IC50=4000nM
    43400 1rfj:A (2.0) BS01 CA ? GO:0005509 ... Q7DMN9 15213382

    Reference:
  • Chengxin Zhang, Xi Zhang, Peter L Freddolino, and Yang Zhang. BioLiP2: an updated structure database for biologically relevent ligand-protein interactions, Nucleic Acids Research, gkad630 (2023).
  • Jianyi Yang, Ambrish Roy, and Yang Zhang. BioLiP: a semi-manually curated database for biologically relevant ligand-protein interactions, Nucleic Acids Research, 41: D1096-D1103 (2013) (download the PDF file).
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