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●TM-score ●TM-align ●US-align ●MM-align ●RNA-align ●NW-align ●LS-align ●EDTSurf ●MVP ●MVP-Fit ●SPICKER ●HAAD ●PSSpred ●3DRobot ●MR-REX ●I-TASSER-MR ●SVMSEQ ●NeBcon ●ResPRE ●TripletRes ●DeepPotential ●WDL-RF ●ATPbind ●DockRMSD ●DeepMSA ●FASPR ●EM-Refiner ●GPU-I-TASSER

●BioLiP ●E. coli ●GLASS ●GPCR-HGmod ●GPCR-RD ●GPCR-EXP ●Tara-3D ●TM-fold ●DECOYS ●POTENTIAL ●RW/RWplus ●EvoEF ●HPSF ●THE-DB ●ADDRESS ●Alpaca-Antibody ●CASP7 ●CASP8 ●CASP9 ●CASP10 ●CASP11 ●CASP12 ●CASP13 ●CASP14

BioLiP
Download all results in tab-seperated text for 197122 receptor-ligand interactions, whose format is explained at readme.txt.
  • Hover over PDB to view the title of the structure. Click PDB to view the structure at the RCSB PDB database. Resolution -1.00 means the resolution is unavailable, e.g., for NMR structures.
  • Click Site # to view the binding site structure. Hover over Site # to view the binding residues.
  • Hover over Ligand to view the full ligand name. Click Ligand to view the 2D diagram and other detail information of the ligand.
  • Hover over EC number to view the full name of enzymatic activity.
  • Hover over GO terms to view all GO terms.
  • Hover over UniProt to view the protein name.

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    # PDB
    (Resolution Å)     		Site # Ligand EC number GO terms UniProt PubMed Binding
    affinity
    160601 7q7f:L (2.75) BS07 FE ? GO:0009772 ... P0C0Y8 N/A
    160602 7q7f:M (2.75) BS07 FE ? GO:0009772 ... P0C0Y9 N/A
    160603 7q7g:L (2.69) BS06 FE ? GO:0009772 ... P0C0Y8 N/A
    160604 7q7g:M (2.69) BS07 FE ? GO:0009772 ... P0C0Y9 N/A
    160605 7q7h:L (2.49) BS07 FE ? GO:0009772 ... P0C0Y8 N/A
    160606 7q7h:M (2.49) BS07 FE ? GO:0009772 ... P0C0Y9 N/A
    160607 7q7j:L (2.69) BS07 FE ? GO:0009772 ... P0C0Y8 N/A
    160608 7q7j:M (2.69) BS07 FE ? GO:0009772 ... P0C0Y9 N/A
    160609 7q7m:L (2.55) BS07 FE ? GO:0009772 ... P0C0Y8 N/A
    160610 7q7m:M (2.55) BS07 FE ? GO:0009772 ... P0C0Y9 N/A
    160611 7q7n:L (2.87) BS07 FE ? GO:0009772 ... P0C0Y8 N/A
    160612 7q7n:M (2.87) BS07 FE ? GO:0009772 ... P0C0Y9 N/A
    160613 7q7o:L (2.65) BS07 FE ? GO:0009772 ... P0C0Y8 N/A
    160614 7q7o:M (2.65) BS07 FE ? GO:0009772 ... P0C0Y9 N/A
    160615 7q7p:LLL (2.4) BS06 FE2 ? GO:0009772 ... P06009 35647917
    160616 7q7p:MMM (2.4) BS04 FE2 ? GO:0009772 ... P06010 35647917
    160617 7q7q:LLL (2.25) BS05 FE2 ? GO:0009772 ... P06009 35647917
    160618 7q7q:MMM (2.25) BS03 FE2 ? GO:0009772 ... P06010 35647917
    160619 7q87:A (1.73) BS01 ZN 3.4.17.18 GO:0004181 ... P29068 N/A
    160620 7q87:A (1.73) BS04 CA 3.4.17.18 GO:0004181 ... P29068 N/A
    160621 7q87:A (1.73) BS05 CA 3.4.17.18 GO:0004181 ... P29068 N/A
    160622 7q87:A (1.73) BS06 CA 3.4.17.18 GO:0004181 ... P29068 N/A
    160623 7q87:A (1.73) BS07 CA 3.4.17.18 GO:0004181 ... P29068 N/A
    160624 7q87:A (1.73) BS08 CA 3.4.17.18 GO:0004181 ... P29068 N/A
    160625 7q87:A (1.73) BS09 CA 3.4.17.18 GO:0004181 ... P29068 N/A
    160626 7q8e:A (2.90002) BS01 ZN 6.3.5.13 GO:0005524 ... A0A0H2WZQ7 N/A
    160627 7q8e:C (2.90002) BS01 ZN 6.3.5.13 GO:0005524 ... A0A0H2WZQ7 N/A
    160628 7q9e:B (1.7) BS02 CO 1.14.14.- GO:0004497 ... D5DF35 36151590
    160629 7qan:AAA (2.01) BS02 MG ? GO:0004497 ... F4F6Q5 36314667
    160630 7qan:BBB (2.01) BS02 MG ? GO:0004497 ... F4F6Q5 36314667
    160631 7qan:BBB (2.01) BS03 MG ? GO:0004497 ... F4F6Q5 36314667
    160632 7qaz:A (2.11) BS01 CO ? N/A H2J4R1 35508653
    160633 7qaz:B (2.11) BS02 CO ? N/A H2J4R1 35508653
    160634 7qaz:B (2.11) BS03 CO ? N/A H2J4R1 35508653
    160635 7qaz:C (2.11) BS01 CO ? N/A H2J4R1 35508653
    160636 7qaz:C (2.11) BS02 CO ? N/A H2J4R1 35508653
    160637 7qb0:A (-1.00) BS01 ZN ? N/A P49023 N/A
    160638 7qb0:A (-1.00) BS02 ZN ? N/A P49023 N/A
    160639 7qb0:A (-1.00) BS03 ZN ? N/A P49023 N/A
    160640 7qb0:A (-1.00) BS04 ZN ? N/A P49023 N/A
    160641 7qb5:111 (1.728) BS02 CA 2.7.7.48
    3.4.22.28
    3.4.22.29
    3.6.1.15    		 GO:0005198 ... V9VEF3 35425270
    160642 7qb5:111 (1.728) BS03 CA 2.7.7.48
    3.4.22.28
    3.4.22.29
    3.6.1.15    		 GO:0005198 ... V9VEF3 35425270
    160643 7qb5:444 (1.728) BS01 CA 2.7.7.48
    3.4.22.28
    3.4.22.29
    3.6.1.15    		 GO:0005198 ... V9VEF3 35425270
    160644 7qba:B (3.78) BS01 MG ? GO:0005524 ... P19844 35896743
    160645 7qba:C (3.78) BS02 MG ? GO:0005524 ... P19844 35896743
    160646 7qba:H (3.78) BS01 CA 1.7.2.4 GO:0004129 ... P19573 35896743
    160647 7qba:I (3.78) BS01 CA 1.7.2.4 GO:0004129 ... P19573 35896743
    160648 7qbd:A (4.18) BS02 CA ? GO:0005515 ... P20062 35218573
    160649 7qbd:B (4.18) BS02 CA ? GO:0005515 ... P20062 35218573
    160650 7qbd:C (4.18) BS01 CA ? N/A Q9NPF0 35218573
    160651 7qbd:C (4.18) BS02 CA ? N/A Q9NPF0 35218573
    160652 7qbd:D (4.18) BS01 CA ? N/A Q9NPF0 35218573
    160653 7qbd:D (4.18) BS02 CA ? N/A Q9NPF0 35218573
    160654 7qbe:B (3.0) BS01 CA ? N/A Q9NPF0 35218573
    160655 7qbe:B (3.0) BS02 CA ? N/A Q9NPF0 35218573
    160656 7qbe:D (3.0) BS01 CA ? N/A Q9NPF0 35218573
    160657 7qbe:D (3.0) BS02 CA ? N/A Q9NPF0 35218573
    160658 7qbf:C (1.85) BS01 CA ? N/A Q9NPF0 35218573
    160659 7qbf:C (1.85) BS02 CA ? N/A Q9NPF0 35218573
    160660 7qbg:B (2.69) BS01 CA ? N/A Q9NPF0 35218573
    160661 7qbg:B (2.69) BS02 CA ? N/A Q9NPF0 35218573
    160662 7qbg:D (2.69) BS01 CA ? N/A Q9NPF0 35218573
    160663 7qbg:D (2.69) BS02 CA ? N/A Q9NPF0 35218573
    160664 7qbg:E (2.69) BS01 CA N/A N/A N/A 35218573
    160665 7qbg:G (2.69) BS01 CA N/A N/A N/A 35218573
    160666 7qbh:AAA (1.223) BS01 ZN 4.2.1.1
    4.2.1.69    		 GO:0002009 ... P00918 35238480
    160667 7qbk:A (2.26) BS01 MN3 ? GO:0009263 ... Q4J9R6 35175627
    160668 7qbk:A (2.26) BS02 FE ? GO:0009263 ... Q4J9R6 35175627
    160669 7qbp:A (1.38) BS01 MN3 1.-.-.- GO:0009263 ... A4F7B2 35175627
    160670 7qbp:A (1.38) BS02 FE 1.-.-.- GO:0009263 ... A4F7B2 35175627
    160671 7qbs:A (2.327) BS03 FE 2.1.1.379 GO:0003824 ... Q8THG6 35110733
    160672 7qbt:A (1.9) BS02 FE 2.1.1.379 GO:0003824 ... Q8THG6 35110733
    160673 7qbt:B (1.9) BS02 FE 2.1.1.379 GO:0003824 ... Q8THG6 35110733
    160674 7qbt:C (1.9) BS02 FE 2.1.1.379 GO:0003824 ... Q8THG6 35110733
    160675 7qbt:D (1.9) BS02 FE 2.1.1.379 GO:0003824 ... Q8THG6 35110733
    160676 7qbu:A (2.298) BS02 FE 2.1.1.379 GO:0003824 ... Q8THG6 35110733
    160677 7qbu:B (2.298) BS02 FE 2.1.1.379 GO:0003824 ... Q8THG6 35110733
    160678 7qbv:A (2.701) BS02 FE 2.1.1.379 GO:0003824 ... Q8THG6 35110733
    160679 7qbv:B (2.701) BS02 FE 2.1.1.379 GO:0003824 ... Q8THG6 35110733
    160680 7qbv:C (2.701) BS02 FE 2.1.1.379 GO:0003824 ... Q8THG6 35110733
    160681 7qbv:D (2.701) BS02 FE 2.1.1.379 GO:0003824 ... Q8THG6 35110733
    160682 7qbz:A (3.25) BS02 MG N/A GO:0000166 ... N/A 34951590
    160683 7qc0:A (3.11) BS02 MG N/A GO:0000166 ... N/A 34951590
    160684 7qc8:A (1.8) BS01 ZN 4.3.2.10 GO:0000105 ... Q9X0C6 35841720
    160685 7qca:LGG (2.79) BS02 ZN ? GO:0003735 ... S7W7C6 N/A
    160686 7qca:LJJ (2.79) BS02 ZN N/A GO:0003735 ... N/A N/A
    160687 7qca:LMM (2.79) BS02 ZN ? GO:0003735 ... S7XSQ3 N/A
    160688 7qca:LOO (2.79) BS02 ZN ? GO:0003735 ... S7XL20 N/A
    160689 7qca:LPP (2.79) BS02 ZN N/A GO:0003735 ... N/A N/A
    160690 7qca:SAA (2.79) BS02 ZN ? GO:0003729 ... S7W9M7 N/A
    160691 7qca:SBB (2.79) BS02 ZN N/A GO:0003735 ... N/A N/A
    160692 7qca:SDD (2.79) BS02 ZN ? GO:0003735 ... S7XJB1 N/A
    160693 7qca:SFF (2.79) BS02 ZN ? GO:0003735 ... S7W9X5 N/A
    160694 7qcd:C (8.0) BS01 ZN 2.3.2.- GO:0000724 ... P38632 35993814
    160695 7qcd:D (8.0) BS01 ZN 2.3.2.27 GO:0000724 ... Q07913 35993814
    160696 7qcd:D (8.0) BS02 ZN 2.3.2.27 GO:0000724 ... Q07913 35993814
    160697 7qce:A (2.1) BS01 ZN ? GO:0010048 ... A0A8B7MWH5 36174674
    160698 7qce:A (2.1) BS02 ZN ? GO:0010048 ... A0A8B7MWH5 36174674
    160699 7qce:A (2.1) BS03 ZN ? GO:0010048 ... A0A8B7MWH5 36174674
    160700 7qce:B (2.1) BS01 ZN ? GO:0010048 ... A0A8B7MWH5 36174674
    160701 7qce:B (2.1) BS02 ZN ? GO:0010048 ... A0A8B7MWH5 36174674
    160702 7qcg:A (1.75) BS02 ZN 2.1.1.56
    2.1.1.57
    2.7.7.48
    2.7.7.50
    3.1.13.-
    3.4.19.12
    3.4.22.-
    3.4.22.69
    3.6.4.12
    3.6.4.13
    4.6.1.-    		 GO:0006508 ... P0DTD1 35480391
    160703 7qch:A (1.88) BS02 ZN 2.1.1.56
    2.1.1.57
    2.7.7.48
    2.7.7.50
    3.1.13.-
    3.4.19.12
    3.4.22.-
    3.4.22.69
    3.6.4.12
    3.6.4.13
    4.6.1.-    		 GO:0006508 ... P0DTD1 35480391
    160704 7qci:A (1.76) BS02 ZN 2.1.1.56
    2.1.1.57
    2.7.7.48
    2.7.7.50
    3.1.13.-
    3.4.19.12
    3.4.22.-
    3.4.22.69
    3.6.4.12
    3.6.4.13
    4.6.1.-    		 GO:0006508 ... P0DTD1 35480391
    160705 7qcj:A (1.84) BS02 ZN 2.1.1.56
    2.1.1.57
    2.7.7.48
    2.7.7.50
    3.1.13.-
    3.4.19.12
    3.4.22.-
    3.4.22.69
    3.6.4.12
    3.6.4.13
    4.6.1.-    		 GO:0006508 ... P0DTD1 35480391
    160706 7qck:A (1.92) BS02 ZN 2.1.1.56
    2.1.1.57
    2.7.7.48
    2.7.7.50
    3.1.13.-
    3.4.19.12
    3.4.22.-
    3.4.22.69
    3.6.4.12
    3.6.4.13
    4.6.1.-    		 GO:0006508 ... P0DTD1 35480391
    160707 7qcl:A (3.36) BS04 CA ? N/A Q02817 37031240
    160708 7qcl:B (3.36) BS04 CA ? N/A Q02817 37031240
    160709 7qcm:A (1.77) BS02 ZN 2.1.1.56
    2.1.1.57
    2.7.7.48
    2.7.7.50
    3.1.13.-
    3.4.19.12
    3.4.22.-
    3.4.22.69
    3.6.4.12
    3.6.4.13
    4.6.1.-    		 GO:0006508 ... P0DTD1 35480391
    160710 7qcn:A (3.4) BS01 CA ? N/A Q02817 37031240
    160711 7qcn:B (3.4) BS01 CA ? N/A Q02817 37031240
    160712 7qcu:A (3.25) BS01 CA ? N/A Q02817 37031240
    160713 7qcu:B (3.25) BS01 CA ? N/A Q02817 37031240
    160714 7qdn:A (1.695) BS03 MG 2.7.1.40 GO:0000287 ... P30613 35290845
    160715 7qdn:B (1.695) BS03 MG 2.7.1.40 GO:0000287 ... P30613 35290845
    160716 7qdn:C (1.695) BS03 MG 2.7.1.40 GO:0000287 ... P30613 35290845
    160717 7qdn:D (1.695) BS03 MG 2.7.1.40 GO:0000287 ... P30613 35290845
    160718 7qdn:E (1.695) BS03 MG 2.7.1.40 GO:0000287 ... P30613 35290845
    160719 7qdn:F (1.695) BS03 MG 2.7.1.40 GO:0000287 ... P30613 35290845
    160720 7qdn:G (1.695) BS03 MG 2.7.1.40 GO:0000287 ... P30613 35290845
    160721 7qdn:H (1.695) BS03 MG 2.7.1.40 GO:0000287 ... P30613 35290845
    160722 7qe1:A (1.95) BS01 ZN N/A GO:0004553 ... N/A 35799064
    160723 7qe1:A (1.95) BS02 ZN N/A GO:0004553 ... N/A 35799064
    160724 7qe1:A (1.95) BS03 ZN N/A GO:0004553 ... N/A 35799064
    160725 7qe1:B (1.95) BS01 ZN N/A GO:0004553 ... N/A 35799064
    160726 7qe1:B (1.95) BS02 ZN N/A GO:0004553 ... N/A 35799064
    160727 7qe1:B (1.95) BS03 ZN N/A GO:0004553 ... N/A 35799064
    160728 7qe1:B (1.95) BS04 ZN N/A GO:0004553 ... N/A 35799064
    160729 7qe1:B (1.95) BS05 ZN N/A GO:0004553 ... N/A 35799064
    160730 7qe2:A (2.15) BS02 ZN N/A GO:0004553 ... N/A 35799064
    160731 7qe2:A (2.15) BS03 ZN N/A GO:0004553 ... N/A 35799064
    160732 7qe2:A (2.15) BS04 ZN N/A GO:0004553 ... N/A 35799064
    160733 7qe2:A (2.15) BS05 ZN N/A GO:0004553 ... N/A 35799064
    160734 7qe2:B (2.15) BS02 ZN N/A GO:0004553 ... N/A 35799064
    160735 7qe2:B (2.15) BS03 ZN N/A GO:0004553 ... N/A 35799064
    160736 7qe2:B (2.15) BS04 ZN N/A GO:0004553 ... N/A 35799064
    160737 7qe7:C () BS01 ZN ? N/A N/A N/A
    160738 7qe7:C () BS02 ZN ? N/A N/A N/A
    160739 7qe7:C () BS03 ZN ? N/A N/A N/A
    160740 7qe7:N () BS01 ZN ? N/A N/A N/A
    160741 7qe7:S () BS01 ZN ? N/A N/A N/A
    160742 7qe7:S () BS02 ZN ? N/A N/A N/A
    160743 7qea:A (2.28) BS02 ZN N/A GO:0004553 ... N/A 35799064
    160744 7qea:A (2.28) BS03 ZN N/A GO:0004553 ... N/A 35799064
    160745 7qea:A (2.28) BS04 ZN N/A GO:0004553 ... N/A 35799064
    160746 7qea:A (2.28) BS05 ZN N/A GO:0004553 ... N/A 35799064
    160747 7qea:A (2.28) BS06 ZN N/A GO:0004553 ... N/A 35799064
    160748 7qea:B (2.28) BS02 ZN N/A GO:0004553 ... N/A 35799064
    160749 7qea:B (2.28) BS03 ZN N/A GO:0004553 ... N/A 35799064
    160750 7qea:B (2.28) BS04 ZN N/A GO:0004553 ... N/A 35799064
    160751 7qea:B (2.28) BS05 ZN N/A GO:0004553 ... N/A 35799064
    160752 7qee:A (2.374) BS01 ZN N/A GO:0004553 ... N/A 35799064
    160753 7qee:A (2.374) BS02 ZN N/A GO:0004553 ... N/A 35799064
    160754 7qee:A (2.374) BS03 ZN N/A GO:0004553 ... N/A 35799064
    160755 7qee:A (2.374) BS04 ZN N/A GO:0004553 ... N/A 35799064
    160756 7qee:B (2.374) BS01 ZN N/A GO:0004553 ... N/A 35799064
    160757 7qee:B (2.374) BS02 ZN N/A GO:0004553 ... N/A 35799064
    160758 7qee:B (2.374) BS03 ZN N/A GO:0004553 ... N/A 35799064
    160759 7qee:B (2.374) BS04 ZN N/A GO:0004553 ... N/A 35799064
    160760 7qee:B (2.374) BS05 ZN N/A GO:0004553 ... N/A 35799064
    160761 7qef:A (2.41) BS02 ZN N/A GO:0004553 ... N/A 35799064
    160762 7qef:A (2.41) BS03 ZN N/A GO:0004553 ... N/A 35799064
    160763 7qef:A (2.41) BS04 ZN N/A GO:0004553 ... N/A 35799064
    160764 7qef:A (2.41) BS05 ZN N/A GO:0004553 ... N/A 35799064
    160765 7qef:A (2.41) BS06 ZN N/A GO:0004553 ... N/A 35799064
    160766 7qef:B (2.41) BS02 ZN N/A GO:0004553 ... N/A 35799064
    160767 7qef:B (2.41) BS03 ZN N/A GO:0004553 ... N/A 35799064
    160768 7qef:B (2.41) BS04 ZN N/A GO:0004553 ... N/A 35799064
    160769 7qef:B (2.41) BS05 ZN N/A GO:0004553 ... N/A 35799064
    160770 7qef:B (2.41) BS06 ZN N/A GO:0004553 ... N/A 35799064
    160771 7qej:A (1.81) BS02 MG ? N/A P0DV33 36602305
    160772 7qen:A (3.46) BS05 MG ? GO:0000070 ... P38989 35653469
    160773 7qen:B (3.46) BS03 MG ? GO:0000070 ... Q12267 35653469
    160774 7qep:D6 (2.7) BS02 ZN ? GO:0003729 ... Q8SRN2 N/A
    160775 7qep:D7 (2.7) BS02 ZN ? GO:0000028 ... Q8SS34 N/A
    160776 7qep:O7 (2.7) BS02 ZN ? GO:0003723 ... Q8SRH5 N/A
    160777 7qep:P0 (2.7) BS02 ZN ? GO:0003735 ... Q8SRH8 N/A
    160778 7qep:P2 (2.7) BS02 ZN ? GO:0003735 ... Q8SR18 N/A
    160779 7qep:P3 (2.7) BS02 ZN ? GO:0003735 ... Q8SRJ2 N/A
    160780 7qfb:A (2.05) BS02 MN 3.1.3.16 GO:0016787 ... P62136 36261419
    160781 7qfb:A (2.05) BS03 MN 3.1.3.16 GO:0016787 ... P62136 36261419
    160782 7qfi:B (1.5) BS01 CA ? N/A Q5FLN0 38838019
    160783 7qfy:A (1.62) BS01 ZN 3.4.24.- GO:0004222 ... M1UZ70 N/A
    160784 7qfy:A (1.62) BS02 CA 3.4.24.- GO:0004222 ... M1UZ70 N/A
    160785 7qfy:A (1.62) BS03 CA 3.4.24.- GO:0004222 ... M1UZ70 N/A
    160786 7qg4:A (2.08) BS01 ZN N/A GO:0004553 ... N/A 35799064
    160787 7qg4:A (2.08) BS02 ZN N/A GO:0004553 ... N/A 35799064
    160788 7qg4:A (2.08) BS03 ZN N/A GO:0004553 ... N/A 35799064
    160789 7qg4:A (2.08) BS04 ZN N/A GO:0004553 ... N/A 35799064
    160790 7qg4:A (2.08) BS05 ZN N/A GO:0004553 ... N/A 35799064
    160791 7qg4:A (2.08) BS06 ZN N/A GO:0004553 ... N/A 35799064
    160792 7qg4:B (2.08) BS01 ZN N/A GO:0004553 ... N/A 35799064
    160793 7qg4:B (2.08) BS02 ZN N/A GO:0004553 ... N/A 35799064
    160794 7qg4:B (2.08) BS03 ZN N/A GO:0004553 ... N/A 35799064
    160795 7qga:A (1.5) BS01 ZN 3.1.3.35
    3.1.3.5
    3.1.3.89
    3.1.3.91
    3.1.3.99    		 GO:0000166 ... P21589 35080883
    160796 7qga:A (1.5) BS02 ZN 3.1.3.35
    3.1.3.5
    3.1.3.89
    3.1.3.91
    3.1.3.99    		 GO:0000166 ... P21589 35080883
    160797 7qga:A (1.5) BS03 CA 3.1.3.35
    3.1.3.5
    3.1.3.89
    3.1.3.91
    3.1.3.99    		 GO:0000166 ... P21589 35080883
    160798 7qgg:SX (2.86) BS02 MG ? GO:0003735 ... A0A340WVW7 36038000
    160799 7qgg:Sa (2.86) BS03 ZN ? GO:0002181 ... A0A2K5LQY6 36038000
    160800 7qgg:Sf (2.86) BS02 ZN ? GO:0003735 ... P62982 36038000
    Reference:
  • Chengxin Zhang, Xi Zhang, Peter L Freddolino, and Yang Zhang. BioLiP2: an updated structure database for biologically relevent ligand-protein interactions, Nucleic Acids Research, gkad630 (2023).
  • Jianyi Yang, Ambrish Roy, and Yang Zhang. BioLiP: a semi-manually curated database for biologically relevant ligand-protein interactions, Nucleic Acids Research, 41: D1096-D1103 (2013) (download the PDF file).

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